2012.12.08 06:17 WinterAnomaly Imaginary Maps - Your source for fictional maps.
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2024.05.14 18:08 Mophandel Archaeotherium, the King of the White River Badlands
Art by Bob Nicholls submitted by Mophandel to badassanimals [link] [comments] Nowadays, when we envision the words “prey,” among modern mammalian fauna, few taxa come to mind as quickly as the hoofed mammals, better known as the ungulates. Indeed, for the better part of their entire evolutionary history, the ungulates have become entirely indistinguishable from the term “prey.” Across their two major modern branches, the artiodactyls (the “even-toed ungulates,” such as bovids, pigs, deer, hippos and giraffes) and the perissodactyls (the “odd-toed ungulates,” including horses, rhinos and tapir), the ungulates too have created an empire spanning nearly every continent, establishing themselves as the the dominant herbivores throughout their entire range. However, as a price for such success, their lot as herbivores have forced them into an unenviable position: being the food for the predators. Indeed, throughout the diets of most modern predators, ungulates make up the majority, if not the entirety, of their diet, becoming their counterparts in this evolutionary dance of theirs. They have become the lamb to their wolf, the zebra to their lion, the stag to their tiger. If there is a predator in need of lunch, chances are that there is an ungulate there to provide it. Of course, such a dynamic is not necessarily a recent innovation. For the last 15-20 million years, across much of the world, both new and old, the ungulates have served as prey for these predators through it all. Over the course of whole epochs, these two groups have played into these roles for millions of years, coevolving with each other in an eons-long game of cat-and-mouse. The shoes they fill are not new, but have existed for ages, and within their niches they have cultivated their roles to perfection. Indeed, with such a tenured history, it seems hardly surprising the ungulates are wholly inseparable from the terms “prey,” itself. However, while this is the case now, as it has been for the last 15-20 million years, go back far enough, and we see that this dynamic is not as set in stone as we would think. Indeed, back during the Eocene and Oligocene, during the very earliest days of age of mammals, things were very different for the ungulates. While today they are considered little more than food for modern predators, during these olden days, the ungulates weren’t quite so benign. In fact, far from being fodder for top predators, the ungulates had turned the tables, instead becoming top predators themselves. Indeed, though nearly unheard of today, throughout much of the Eocene and Oligocene, carnivorous ungulates thrived in abundance, developing specializations for catching large prey and establishing themselves as top predators that competed alongside the more traditional carnivores, and even dominating them in some instances. Given such success, it’s no wonder that multiple such clades had arisen during this time. Such predators included the arctocyonids, a lineage of (ironically) hoof-less ungulates with large jaws and sharp teeth for capturing large prey. There were also the mesonychians, a lineage of dog-like ungulates with massive skulls and jaws that allowed them to reign as the top predator across much of the Eocene. However, among these various lineages, one stands stands out among the rest, by far. Arising during the Eocene, this lineage, though superficially resembling modern pigs, hailed from one an ancient lineage of artiodactyls far removed from swine or most other ungulates in general, with few close relatives alive today. Through perhaps not the most predatory of the bunch, it was among the most formidable, as their superficially pig-like appearance came with giant predatory jaws and teeth unlike anything from the modern era. And of course, as if all of that wasn’t enough, this lineage also went on to earn arguably one of the most badass nicknames of any lineage of mammals, period. These predators, of course, were the entelodonts, a.k.a the “hell-pigs.” More so than any other predatory ungulate lineage, these formidable ungulates were the ones to turn the current paradigm upside down, becoming some of the largest and most dominant carnivores in their landscape, even with (and often in spite of) the presence of more traditional predators. Through impressive size, fearsome teeth and sheer tenacity, these animals became the top dogs of their time, ruling as behemoth-kings of their Paleogene kingdoms, domineering all comers, and throughout the ranks, one entelodont in particular demonstrated such dominance the best. Though not the largest or most powerful of their kind, it is one of the most iconic, being among the most well-known members of its lineage to date. Moreover, this enteledont also has some of the most complete life histories ever seen out of this clade, with its brutality and predatory prowess being displayed in the fossil record in a way seen in no other member of its kind. More than anything else, however, it was this predator that best turned the notion of “ungulates being prey” on its head, living in an environment that bore some of the largest carnivoran hypercarnivores to date and still reigning as the undisputed top predator of its domain. This fearsome beast was none other than Archaeotherium, icon of the entelodonts, terror of the Oligocene American west and undisputed king of the White River badlands. The rise of Archaeotherium (and of entelodonts in general) is closely tied to the ascendancy of carnivorous ungulates as a whole, one of the earliest evolutionary success stories of the entire Cenozoic. Having become their own derived clade since the late Cretaceous, the ungulates were remarkably successful during the early Paleogene, as they were among the first mammalian clades to reach large sizes during those early days after the non-avian dinosaurs had gone extinct. As such, it was with incredible swiftness that, as the Paleogene progressed, the ungulates swooped upon the various niches left empty by the K-Pg mass extinction that killed the dinosaurs. This of course included the herbivorous niches we would know them for today, but this also included other, much more carnivore roles. Indeed, early on during the Paleogene, it was the ungulates that first seized the roles of large mammalian predators, becoming some the earliest large mammalian carnivores to ever live, well before even the carnivorans. Such predators included the arctocyonids, a lineage of vaguely dog-like, hoof-less ungulates with robust jaws and sharpened teeth that acted as some of earliest large carnivores of the Paleocene, with genera such as Arctocyon mumak getting up to the size of big cats. Even more prolific were the mesonychids. More so than what pretty much any other lineage of predator, it was the mesonychids that would stand out as the earliest dominant predators of the early Cenozoic. Growing up to the size of bears and with enormous, bone-crushing jaws, the mesonychids were among the most powerful and successful predators on the market at that time, with a near-global range and being capable of subjugating just about any other predator in their environments. Indeed, they, along with other carnivorous ungulates (as well as ungulates in general), were experiencing a golden age during this time, easily being the most prolific predators of the age. Given such prevalence, it should be no surprise that there would be yet another lineage of predatory ungulates would throw their hat into the ring, and by early Eocene, that contender would none other than the entelodonts. The very first entelodonts had arisen from artiodactyl ancestors during the Eocene epoch, at a time when artiodactyls were far more diverse and bizarre than they are now. Through today known from their modern herbivorous representatives such as bovines, deer, and antelope, during the Paleocene and Eocene, the artiodacyls, as with most ungulates of that time, were stronger and far more predaceous, particularly when it came to one such clade of artiodactyls, the cetacodontamorphs. Only known today from hippos and another group of artiodactyls (one which will become relevant later), the cetacodantomorphs emerged out of Asia around 55 million years ago, at around the same time that artiodactyls themselves had made their debut. These animals included the first truly predatory artiodactyls, with many of them possessing large skulls with powerful jaws and sharp, predatory teeth. Among their ranks included animals as puny as Indohyus, a piscivorous artiodactyl the size of a cat, to as formidable as Andrewsarchus, a giant, bison-sized predator often touted as one of the largest predatory mammals to ever live. Given such a predatory disposition, it wouldn’t be long until this clade produced a lineage of truly diverse, truly successful predators, and by around 40 million years ago, that is exactly what they did, as it was at that time that the entelodonts themselves first emerged. From their Asian homeland, the entelodonts spread across the world, spreading through not only most of Eurasia but also colonizing North America as well, with genera such as Brachyhyops being found across both continents. Here, in this North American frontier, the entelodonts began to diversify further, turning into their most successful and formidable forms yet, and it was around the late Eocene and early Oligocene that Archaeotherium itself had entered the scene. Just from a passing glance at Archaeotherium, it is clear how exactly it (as well as the other entelodonts) earned the nickname of “hell-pigs.” It was a bruiser for starters; its body bore a robust, pig-like physique, with prominent neural spines and their associated musculature forming a hump around the shoulder region, similar to the hump of a bison. With such a bulky physique came with it impressive size; the average A. mortoni had a head-body length of roughly 1.6-2.0 m (5.3-6.6 ft), a shoulder height of 1.2 m (4 ft) and a body mass of around 180 kg (396 lb) in weight (Boardman & Secord, 2013; Joeckel, 1990). At such sizes, an adult Archaeotherium the size of a large male black bear. However, they had the potential to get even bigger. While most Archaeotherium specimens were around the size described above, a select few specimens, labeled under the synonymous genus “Megachoerus,” are found to be much larger, with skulls getting up to 66% longer than average A. mortoni specimens (Foss, 2001; Joeckel, 1990). At such sizes and using isometric scaling, such massive Archaeotherium specimens would attained body lengths over 2.5 m (8.2 ft) and would have reached weighs well over 500 kg (1100 lb), or as big as a mature male polar bear. Indeed, at such sizes, it is already abundantly evident that Archaeotherium is a force to be recorded with. However, there was more to these formidable animals than sheer size alone. Behind all that bulk was an astoundingly swift and graceful predator, especially in terms of locomotion. Indeed, the hoofed feet of Archaeotherium, along with other entelodonts, sported several adaptations that gave it incredible locomotive efficiency, essentially turning it into a speed demon of the badlands. Such adaptations include longer distal leg elements (e.g. the radius and tibia) than their proximal counterparts (e.g. the humerus and femur), fusion of the radius and ulna for increased running efficiency, the loss of the clavicle (collar-bone) to allow for greater leg length, the loss of the acromion to enhance leg movement along the fore-and-aft plane, the loss of digits to reduce the mass of the forelimb, the fusion of the ectocuneiform and the mesocuneiform wrist-bones, among many other such traits (Theodore, 1996) . Perhaps most significant of these adaptations is the evolution of the “double-pulley astragalus (ankle-bone),” a specialized modification of the ankle that, while restricting rotation and side-to-side movement at the ankle-joint, allows for greater rotation in the fore-and-aft direction, thus allowing for more more powerful propulsion from the limbs, faster extension and retraction of the limbs and overall greater locomotive efficiency (Foss, 2001). Of course, such a trait was not only found in entelodonts but in artiodactyls as a whole, likely being a response to predatory pressures from incumbent predatory clades arising at the same time as the artiodactyls (Foss, 2001). However, in the case of the entelodonts, such adaptations were not used for merely escaping predators. Rather, they were used to for another, much more lethal effect… Such notions are further reinforced by the entelodonts most formidable aspect, none either than their fearsome jaws, and in this respect, Archaeotherium excelled. Both for its size and in general, the head of Archaeotherium was massive, measuring 40-50 cm (1.3-1.6 ft) in length among average A. mortoni specimens, to up to 78 cm (~2.6 ft) in the larger “Megachoerus” specimens (Joeckel, 1990). Such massive skulls were supported and supplemented by equally massive neck muscles and ligaments, which attached to massive neural spines on the anterior thoracic vertebrae akin to a bisons hump as well as to the sternum, allowing Archaeotherium to keep its head aloft despite the skulls massive size (Effinger, 1998). Of course, with such a massive skull, it should come as no surprise that such skulls housed exceptionally formidable jaws as well, and indeed, the bite of Archaeotherium was an especially deadly one. Its zygomatic arches (cheek-bones) and its temporal fossa were enlarged and expanded, indicative of massive temporalis muscles that afforded Archaeotherium astoundingly powerful bites (Joeckel, 1990). This is further augmented by Archaeotherium’s massive jugal flanges (bony projections of the cheek), which supported powerful masseter muscles which enhanced chewing and mastication, as well as an enlarged postorbital bar that reinforced the skull against torsional stresses (Foss, 2001). Last but not least, powerful jaws are supplemented by an enlarged gape, facilitated by a low coronoid process and enlarged posterior mandibular tubercles (bony projections originating from the lower jaw), which provided an insertion site for sternum-to-mandible jaw abduction muscles, allowing for a more forceful opening of the jaw (Foss, 2001). All together, such traits suggest a massive and incredibly fearsome bite, perhaps the most formidable of any animal in its environment. Of course, none of such traits are especially indicative of a predatory lifestyle. Indeed, many modern non-predatory ungulates, like hippos, pigs and peccaries, also possess large, formidable skulls and jaws. However, in peeling back the layers, it is found there was more to the skull of Archaeotherium that lies in store. Indeed, when inspecting the animal closely, a unique mosaic of features is revealed; traits that make it out to be much more lethal than the average artiodactyl. On one hand, Archaeotherium possessed many traits similar to those of herbivores animals, as is expected of ungulates. For instance, its jaw musculature that allowed the lower jaw of Archaeotherium a full side-to-side chewing motion as in herbivores (whereas most carnivores can only move their lower jaw up and down)(Effinger, 1998). On the other hand, Archaeotherium wielded many other traits far more lethal in their morphology, less akin to a herbivore and far more akin to a bonafide predator. For instance, the aforementioned enlarged gape of Archaeotherium is a bizarre trait on a supposed herbivore, as such animals do not need large gapes to eat vegetation and thus have smaller, more restricted gapes. Conversely, many predatory lineages have comparatively large gapes, as larger gapes allow for the the jaws to grab on to more effectively larger objects, namely large prey animals (Joeckel, 1990). Such a juxtaposition, however, is most evident when discussing the real killing instruments of Archaeotherium — the teeth. More so than any facet of this animal, the teeth of Archaeotherium are the real stars of the show, showing both how alike it was compared to its herbivores counterparts and more importantly, how it couldn’t be more different. For instance, the molars of Archaeotherium were quite similar to modern herbivores ungulates, in that they were robust, bunodont, and were designed for crushing and grinding, similar in form and function to modern ungulates like peccaries (Joeckel, 1990). However, while the molars give the impression that Archaeotherium was a herbivore, the other teeth tell a very different story. The incisors, for example, were enlarged, sharpened, and fully interlocked (as opposed to the flat-topped incisors seen in herbivores ungulates), creating an incisor array that was seemingly ill-suited for cropping vegetation and much more adept at for gripping, puncturing and cutting (Joeckel, 1990). Even more formidable were the canines. Like the modern pigs from which entelodonts derived their nicknames, the canines of Archaeotherium were sharp and enlarged to form prominent tusk-like teeth, but unlike pigs, they were rounded in cross-section (similar to modern carnivores like big cats, indicating more durable canines that can absorb and resist torsional forces, such as those from struggling prey) and were serrated to form a distinct cutting edge (Effinger, 1998; Joeckel, 1990; Ruff & Van Valkenburgh, 1987). These canines, along with the incisors, interlock to stabilize the jaws while biting and dismantling in a carnivore-like fashion. More strikingly, the canines also seem to act as “occlusal guides,” wherein the canines help align the movement and position of the rear teeth as they come together, allowing for a more efficient shearing action by the rear teeth. This function is seen most prevalently modern carnivorous mammals, and is evidenced by the canine tooth-wear, which is also analogous to modern predators like bears and canids (Joeckel, 1990). Indeed, going off such teeth alone, it is clear that Archaeotherium is far more predatory than expected of an ungulate. However, the real stars of the show, the teeth that truly betray the predatory nature of these ungulates, are the premolars. Perhaps the most carnivore-like teeth in the entelodont’s entire tooth row, the premolars of Archaeotherium, particularly the anterior premolars, are laterally compressed, somewhat conical in shape, and are weakly serrated to bear a cutting edge, giving them a somewhat carnivorous form and function of shearing and slicing (Effinger, 1998). Most strikingly of all, the premolars of Archaeotherium bear unique features similar not to modern herbivores, but to durophagous carnivores like hyenas, particularly apical wear patterns, highly thickened enamel, “zigzag-shaped” enamel prism layers (Hunter-Schraeger bands) on the premolars which is also seen in osteophagous animals like hyenas, and an interlocking premolar interface wherein linear objects (such as bones) inserted into jaws from the side would be pinned between the premolars and crushed (Foss, 2001). Taken together, these features do not suggest a diet of grass or vegetation like other ungulates. Rather, they suggest a far more violent diet, one including flesh as well as hard, durable foods, particularly bone. All in all, the evidence is clear. Archaeotherium and other entelodonts, unlike the rest of their artiodactyl kin, were not the passive herbivores as we envision ungulates today. Rather, they were willing, unrepentant meat-eaters that had a taste for flesh as well as foliage. Of course, even with such lines of evidence, its hard to conclude that Archaeotherium was a true predator. After all, its wide gape and durophagous teeth could have just as easily been used for scavenging or even to eat tough plant matter such as seeds or nuts, as in peccaries and pigs, which themselves share many of the same adaptations as Archaeotherium, include the more carnivorous ones (e.g. the wide gape, using the canines as an occlusal guide, etc.). How exactly do we know that these things were veritable predators and not pretenders to the title. To this end, there is yet one last piece of evidence, one that puts on full display the predatory prowess of Archaeotherium —evidence of a kill itself. Found within oligocene-aged sediment in what is now Wyoming, a collection of various fossil remains was found, each belonging to the ancient sheep-sized camel Poebrotherium, with many of the skeletal remains being disarticulated and even missing whole hindlimbs or even entire rear halves of their body. Tellingly, many of the remains bear extensive bite marks and puncture wounds across their surface. Upon close examination, the spacing and size of the punctures leave only one culprit: Archaeotherium. Of course, such an event could still have been scavenging; the entelodonts were consuming the remains of already dead, decomposed camels, explaining the bite marks. What was far more telling, however, was where the bite marks were found. In addition bite marks being found on the torso and lumbar regions of the camels, various puncture wounds were found on the skull and neck, which were otherwise uneaten. Scavengers rarely feast on the head to begin with; there is very little worthwhile meat on it besides the brain, cheek-muscles and eyes, and even if they did feed on the skull and neck, they would still eat it wholesale, not merely bite it and then leave it otherwise untouched. Indeed, it was clear that this was no mere scavenging event. Rather than merely consuming these camels, Archaeotherium was actively preying upon and killing them, dispatching them via a crushing bite to the skull or neck before dismembering and even bisecting the hapless camels with their powerful jaws to preferentially feast on their hindquarters (likely by swallowing the hindquarters whole, as the pelvis of Poebrotherium was coincidentally the perfect width for Archaeotherium to devour whole), eventually discarding the leftovers in meat caches for later consumption (Sundell, 1999). With this finding, such a feat of brutality leaves no doubt in ones mind as to what the true nature of Archaeotherium was. This was no herbivore, nor was it a simple scavenger. This was an active, rapacious predator, the most powerful in its entire ecosystem. Indeed, with such brutal evidence of predation frozen in time, combined with various dental, cranial, and post cranial adaptations of this formidable animal, it’s possible to paint a picture of how this formidable creature lived. Though an omnivore by trade, willing and able to feast on plant matter such as grass, roots and tubers, Archaeotherium was also a wanton predator that took just about any prey it wanted. Upon detecting its prey, it approached its vicim from ambush before launching itself at blazing speed. From there, its cursorial, hoofed legs, used by other ungulates for escape predation, were here employed to capture prey, carrying it at great speeds as it caught up to its quarry. Having closed the distance with its target, it was then that the entelodont brought its jaws to bear, grabbing hold of the victim with powerful jaws and gripping teeth to bring it to a screeching halt. If the victim is lucky, Archaeotherium will then kill it quickly with a crushing bite to the skull or neck, puncturing the brain or spinal cord and killing its target instantly. If not, the victim is eaten alive, torn apart while it’s still kicking, as modern boars will do today. In any case, incapacitated prey are subsequently dismantled, with the entelodont using its entire head and heavily-muscled necks to bite into and pull apart its victim in devastating “puncture-and pull’ bites (Foss, 2001). Prey would then finally be consumed starting at the hindquarters, with not even the bones of its prey being spared. Such brutality, though far from clean, drove home a singular truth: that during this time, ungulates were not just prey, that they were not the mere “predator-fodder” we know them as today. rather, they themselves were the predators themselves, dominating as superb hunters within their domain and even suppressing clades we know as predators today, least of all the carnivorans. Indeed, during this point in time, the age of the carnivorous ungulates had hit their stride, and more specifically, the age of entelodonts had begun. Of course, more so than any other entelodont, Archaeotherium took to this new age with gusto. Archaeotherium lived from 35-28 million years ago during the late Eocene and early Oligocene in a locality known today as the White River Badlands, a fossil locality nestled along the Great Plains and Rocky Mountains. Though a chalky, barren landscape today, during the time of Archaeotherium, the White River Badlands was a swamp-like floodplain crisscrossed with rivers and interspersed with by a mosaic of forests concentrated around waterways, open woodlands and open plains. As with most ecosystems with such a lush disposition, this locale teemed with life, with ancient hornless rhinos, small horse-like hyracodonts and early camels roaming the open habitats while giant brontotheres, small early horses and strange, sheep-like ungulates called merycoidodonts (also known as “oreodonts”) dwelled within the dense forests. Within this locale, Archaeotherium stalked the open woodlands and riparian forests of its domain. Here, it acted as a dominant predator and scavenger across is territory, filling a niche similar to modern grizzly bears but far more predatory. Among its preferred food items would be plant matter such as roots, foliage and nuts, but also meat in the form of carrion or freshly caught prey. In this respect, smaller ungulates such as the fleet-footed camel Poebrotherium, a known prey item of Archaeotherium, would have made a for choice prey, as its small size would make it easy for Archaeotherium to dispatch with its powerful jaws, while the entelodonts swift legs gave it the speed necessary to keep pace with its agile prey. However, the entelodont didn’t have such a feast all to itself. Just as the badlands teemed with herbivores, so too did it teem with rival predators. Among their ranks included fearsome predators such as Hyaenodon, a powerful, vaguely dog-like predator up to the size of wolves (as in H. horridus) or even lions (as in the Eocene-aged H. megaloides, which was replaced by H. horridus during the Oligocene). Armed with a massive head, fierce jaws and a set of knife-like teeth that could cut down even large prey in seconds, these were some of the most formidable predators on the landscape. There were also the nimravids, cat-like carnivorans that bore saber-teeth to kill large prey in seconds, and included the likes of the lynx-sized Dinictis, the leopard-sized Hoplophoneus and even the jaguar-sized Eusmilus. Furthermore, there were amphicyonids, better known as the bear-dogs. Though known from much larger forms later on in their existence, during the late Eocene and Oligocene, they were much smaller and acted as the “canid-analogues” of the ecosystem, filling a role similar to wolves or coyotes. Last but not least, there were the bathornithid birds, huge cariamiform birds related to modern seriemas but much larger, which filled a niche similar to modern seriemas or secretary birds, albeit on a much larger scale. Given such competition, it would seem that Archaeotherium would have its hands full. However, things are not as they appear. For starters, habitat differences would mitigate high amounts of competition, as both Hyaenodon and the various nimravids occupy more specialized ecological roles (being a plains-specialist and forest-specialist, respectively) than did Archaeotherium, providing a buffer to stave off competition: More importantly, however, none of the aforementioned predators were simply big enough to take Archaeotherium on. During the roughly 7 million years existence of Archaeotherium, the only carnivore that matched it in size was H. megaloides, and even that would have an only applied to average A. mortoni individuals, not to the much larger, bison-sized “Megachoerus” individuals. The next largest predator at that point would be the jaguars-sized Eusmilus (specifically E. adelos) which would have only been a bit more than half the size of even an average A. mortoni. Besides that, virtually every other predator on the landscape was simply outclassed by the much larger entelodont in terms of size and brute strength. As such, within its domain, Archaeotherium had total, unquestioned authority, dominating the other predators in the landscape and likely stealing their kills as well. In fact, just about the only threat Archaeotherium had was other Archaeotherium, as fossil bite marks suggest that this animal regularly and fraglantly engaged in intraspecific combat, usually through face-biting and possibly even jaw-wrestling (Effinger, 1998; Tanke & Currie, 1998). Nevertheless, it was clear that Archaeotherium was the undisputed king of the badlands; in a landscape of hyaenodonts and carnivorans galore, it was a hoofed ungulate that reigned supreme. However, such a reign would not last. As the Eocene transitioned into the Eocene, the planet underwent an abrupt cooling and drying phase known as Eocene-Oligocene Transition or more simply the Grande Coupure. This change in climate would eliminate the sprawling wetlands and river systems that Archaeotherium had been depending on, gradually replacing it with drier and more open habitats. To its credit, Archaeotherium did manage to hang on, persisting well after the Grand-Coupure had taken place, but in the end the damage had been done; Archaeotherium was a dead-man-walking. Eventually, by around 28 million years ago, Archaeotherium would go extinct, perishing due to this change in global climate (Gillham, 2019). Entelodonts as a whole would persist into the Miocene, producing some of their largest forms ever known in the form of the bison-sized Daeodon (which was itself even more carnivorous than Archaeotherium), however they too would meet the same fate as their earlier cousins. By around 15-20 million years ago, entelodonts as a whole would go extinct. However, while the entelodonts may have perished, this was not the end of carnivorous ungulates as a whole. Recall that the cetacodontamorphs, the lineage of artiodactyls that produced the entelodonts, left behind two living descendants. The first among them were the hippos, themselves fairly frequent herbivores. The second of such lineage, however, was a different story. Emerging out of South Asia, this lineage of piscivorous cetacodontamorphs, in a an attempt to further specialize for the fish-hunting lifestyle, began to delve further and further into the water, becoming more and more aquatic and the millennia passed by. At a certain point, these carnivorous artiodactlys had become something completely unrecognizable from their original hoofed forms. Their skin became hairless and their bodies became streamlined for life in water. Their hoofed limbs grew into giant flippers for steering in the water and their previously tiny tails became massive and sported giant tail flukes for aquatic propulsion. Their noses even moved to the tip of their head, becoming a blowhole that would be signature to this clade as a whole. Indeed, this clade was none other than the modern whales, themselves derived, carnivorous ungulates that had specialized for a life in the water, and in doing so, became the some of the most dominant aquatic predators across the globe for millions of years. Indeed, though long gone, the legacy of the entelodonts and of predatory ungulates as a whole, a legacy Archaeotherium itself had helped foster, lives on in these paragons of predatory prowess, showing that the ungulates are more than just the mere “prey” that they are often made out to be. Moreover, given the success that carnivorous ungulates had enjoyed in the past and given how modern omnivorous ungulates like boar dabble in predation themselves, perhaps, in the distant future, this planet may see the rise of carnivorous ungulates once again, following in the footsteps left behind by Archaeotherium and the other predatory ungulates all those millions of years ago. |
2024.05.14 16:25 Mophandel Archaeotherium, the King of the White River Badlands
Art by Bob Nicholls submitted by Mophandel to Naturewasmetal [link] [comments] Nowadays, when we envision the words “prey,” among modern mammalian fauna, few taxa come to mind as quickly as the hoofed mammals, better known as the ungulates. Indeed, for the better part of their entire evolutionary history, the ungulates have become entirely indistinguishable from the term “prey.” Across their two major modern branches, the artiodactyls (the “even-toed ungulates,” such as bovids, pigs, deer, hippos and giraffes) and the perissodactyls (the “odd-toed ungulates,” including horses, rhinos and tapir), the ungulates too have created an empire spanning nearly every continent, establishing themselves as the the dominant herbivores throughout their entire range. However, as a price for such success, their lot as herbivores have forced them into an unenviable position: being the food for the predators. Indeed, throughout the diets of most modern predators, ungulates make up the majority, if not the entirety, of their diet, becoming their counterparts in this evolutionary dance of theirs. They have become the lamb to their wolf, the zebra to their lion, the stag to their tiger. If there is a predator in need of lunch, chances are that there is an ungulate there to provide it. Of course, such a dynamic is not necessarily a recent innovation. For the last 15-20 million years, across much of the world, both new and old, the ungulates have served as prey for these predators through it all. Over the course of whole epochs, these two groups have played into these roles for millions of years, coevolving with each other in an eons-long game of cat-and-mouse. The shoes they fill are not new, but have existed for ages, and within their niches they have cultivated their roles to perfection. Indeed, with such a tenured history, it seems hardly surprising the ungulates are wholly inseparable from the terms “prey,” itself. However, while this is the case now, as it has been for the last 15-20 million years, go back far enough, and we see that this dynamic is not as set in stone as we would think. Indeed, back during the Eocene and Oligocene, during the very earliest days of age of mammals, things were very different for the ungulates. While today they are considered little more than food for modern predators, during these olden days, the ungulates weren’t quite so benign. In fact, far from being fodder for top predators, the ungulates had turned the tables, instead becoming top predators themselves. Indeed, though nearly unheard of today, throughout much of the Eocene and Oligocene, carnivorous ungulates thrived in abundance, developing specializations for catching large prey and establishing themselves as top predators that competed alongside the more traditional carnivores, and even dominating them in some instances. Given such success, it’s no wonder that multiple such clades had arisen during this time. Such predators included the arctocyonids, a lineage of (ironically) hoof-less ungulates with large jaws and sharp teeth for capturing large prey. There were also the mesonychians, a lineage of dog-like ungulates with massive skulls and jaws that allowed them to reign as the top predator across much of the Eocene. However, among these various lineages, one stands stands out among the rest, by far. Arising during the Eocene, this lineage, though superficially resembling modern pigs, hailed from one an ancient lineage of artiodactyls far removed from swine or most other ungulates in general, with few close relatives alive today. Through perhaps not the most predatory of the bunch, it was among the most formidable, as their superficially pig-like appearance came with giant predatory jaws and teeth unlike anything from the modern era. And of course, as if all of that wasn’t enough, this lineage also went on to earn arguably one of the most badass nicknames of any lineage of mammals, period. These predators, of course, were the entelodonts, a.k.a the “hell-pigs.” More so than any other predatory ungulate lineage, these formidable ungulates were the ones to turn the current paradigm upside down, becoming some of the largest and most dominant carnivores in their landscape, even with (and often in spite of) the presence of more traditional predators. Through impressive size, fearsome teeth and sheer tenacity, these animals became the top dogs of their time, ruling as behemoth-kings of their Paleogene kingdoms, domineering all comers, and throughout the ranks, one entelodont in particular demonstrated such dominance the best. Though not the largest or most powerful of their kind, it is one of the most iconic, being among the most well-known members of its lineage to date. Moreover, this enteledont also has some of the most complete life histories ever seen out of this clade, with its brutality and predatory prowess being displayed in the fossil record in a way seen in no other member of its kind. More than anything else, however, it was this predator that best turned the notion of “ungulates being prey” on its head, living in an environment that bore some of the largest carnivoran hypercarnivores to date and still reigning as the undisputed top predator of its domain. This fearsome beast was none other than Archaeotherium, icon of the entelodonts, terror of the Oligocene American west and undisputed king of the White River badlands. The rise of Archaeotherium (and of entelodonts in general) is closely tied to the ascendancy of carnivorous ungulates as a whole, one of the earliest evolutionary success stories of the entire Cenozoic. Having become their own derived clade since the late Cretaceous, the ungulates were remarkably successful during the early Paleogene, as they were among the first mammalian clades to reach large sizes during those early days after the non-avian dinosaurs had gone extinct. As such, it was with incredible swiftness that, as the Paleogene progressed, the ungulates swooped upon the various niches left empty by the K-Pg mass extinction that killed the dinosaurs. This of course included the herbivorous niches we would know them for today, but this also included other, much more carnivore roles. Indeed, early on during the Paleogene, it was the ungulates that first seized the roles of large mammalian predators, becoming some the earliest large mammalian carnivores to ever live, well before even the carnivorans. Such predators included the arctocyonids, a lineage of vaguely dog-like, hoof-less ungulates with robust jaws and sharpened teeth that acted as some of earliest large carnivores of the Paleocene, with genera such as Arctocyon mumak getting up to the size of big cats. Even more prolific were the mesonychids. More so than what pretty much any other lineage of predator, it was the mesonychids that would stand out as the earliest dominant predators of the early Cenozoic. Growing up to the size of bears and with enormous, bone-crushing jaws, the mesonychids were among the most powerful and successful predators on the market at that time, with a near-global range and being capable of subjugating just about any other predator in their environments. Indeed, they, along with other carnivorous ungulates (as well as ungulates in general), were experiencing a golden age during this time, easily being the most prolific predators of the age. Given such prevalence, it should be no surprise that there would be yet another lineage of predatory ungulates would throw their hat into the ring, and by early Eocene, that contender would none other than the entelodonts. The very first entelodonts had arisen from artiodactyl ancestors during the Eocene epoch, at a time when artiodactyls were far more diverse and bizarre than they are now. Through today known from their modern herbivorous representatives such as bovines, deer, and antelope, during the Paleocene and Eocene, the artiodacyls, as with most ungulates of that time, were stronger and far more predaceous, particularly when it came to one such clade of artiodactyls, the cetacodontamorphs. Only known today from hippos and another group of artiodactyls (one which will become relevant later), the cetacodantomorphs emerged out of Asia around 55 million years ago, at around the same time that artiodactyls themselves had made their debut. These animals included the first truly predatory artiodactyls, with many of them possessing large skulls with powerful jaws and sharp, predatory teeth. Among their ranks included animals as puny as Indohyus, a piscivorous artiodactyl the size of a cat, to as formidable as Andrewsarchus, a giant, bison-sized predator often touted as one of the largest predatory mammals to ever live. Given such a predatory disposition, it wouldn’t be long until this clade produced a lineage of truly diverse, truly successful predators, and by around 40 million years ago, that is exactly what they did, as it was at that time that the entelodonts themselves first emerged. From their Asian homeland, the entelodonts spread across the world, spreading through not only most of Eurasia but also colonizing North America as well, with genera such as Brachyhyops being found across both continents. Here, in this North American frontier, the entelodonts began to diversify further, turning into their most successful and formidable forms yet, and it was around the late Eocene and early Oligocene that Archaeotherium itself had entered the scene. Just from a passing glance at Archaeotherium, it is clear how exactly it (as well as the other entelodonts) earned the nickname of “hell-pigs.” It was a bruiser for starters; its body bore a robust, pig-like physique, with prominent neural spines and their associated musculature forming a hump around the shoulder region, similar to the hump of a bison. With such a bulky physique came with it impressive size; the average A. mortoni had a head-body length of roughly 1.6-2.0 m (5.3-6.6 ft), a shoulder height of 1.2 m (4 ft) and a body mass of around 180 kg (396 lb) in weight (Boardman & Secord, 2013; Joeckel, 1990). At such sizes, an adult Archaeotherium the size of a large male black bear. However, they had the potential to get even bigger. While most Archaeotherium specimens were around the size described above, a select few specimens, labeled under the synonymous genus “Megachoerus,” are found to be much larger, with skulls getting up to 66% longer than average A. mortoni specimens (Foss, 2001; Joeckel, 1990). At such sizes and using isometric scaling, such massive Archaeotherium specimens would attained body lengths over 2.5 m (8.2 ft) and would have reached weighs well over 500 kg (1100 lb), or as big as a mature male polar bear. Indeed, at such sizes, it is already abundantly evident that Archaeotherium is a force to be recorded with. However, there was more to these formidable animals than sheer size alone. Behind all that bulk was an astoundingly swift and graceful predator, especially in terms of locomotion. Indeed, the hoofed feet of Archaeotherium, along with other entelodonts, sported several adaptations that gave it incredible locomotive efficiency, essentially turning it into a speed demon of the badlands. Such adaptations include longer distal leg elements (e.g. the radius and tibia) than their proximal counterparts (e.g. the humerus and femur), fusion of the radius and ulna for increased running efficiency, the loss of the clavicle (collar-bone) to allow for greater leg length, the loss of the acromion to enhance leg movement along the fore-and-aft plane, the loss of digits to reduce the mass of the forelimb, the fusion of the ectocuneiform and the mesocuneiform wrist-bones, among many other such traits (Theodore, 1996) . Perhaps most significant of these adaptations is the evolution of the “double-pulley astragalus (ankle-bone),” a specialized modification of the ankle that, while restricting rotation and side-to-side movement at the ankle-joint, allows for greater rotation in the fore-and-aft direction, thus allowing for more more powerful propulsion from the limbs, faster extension and retraction of the limbs and overall greater locomotive efficiency (Foss, 2001). Of course, such a trait was not only found in entelodonts but in artiodactyls as a whole, likely being a response to predatory pressures from incumbent predatory clades arising at the same time as the artiodactyls (Foss, 2001). However, in the case of the entelodonts, such adaptations were not used for merely escaping predators. Rather, they were used to for another, much more lethal effect… Such notions are further reinforced by the entelodonts most formidable aspect, none either than their fearsome jaws, and in this respect, Archaeotherium excelled. Both for its size and in general, the head of Archaeotherium was massive, measuring 40-50 cm (1.3-1.6 ft) in length among average A. mortoni specimens, to up to 78 cm (~2.6 ft) in the larger “Megachoerus” specimens (Joeckel, 1990). Such massive skulls were supported and supplemented by equally massive neck muscles and ligaments, which attached to massive neural spines on the anterior thoracic vertebrae akin to a bisons hump as well as to the sternum, allowing Archaeotherium to keep its head aloft despite the skulls massive size (Effinger, 1998). Of course, with such a massive skull, it should come as no surprise that such skulls housed exceptionally formidable jaws as well, and indeed, the bite of Archaeotherium was an especially deadly one. Its zygomatic arches (cheek-bones) and its temporal fossa were enlarged and expanded, indicative of massive temporalis muscles that afforded Archaeotherium astoundingly powerful bites (Joeckel, 1990). This is further augmented by Archaeotherium’s massive jugal flanges (bony projections of the cheek), which supported powerful masseter muscles which enhanced chewing and mastication, as well as an enlarged postorbital bar that reinforced the skull against torsional stresses (Foss, 2001). Last but not least, powerful jaws are supplemented by an enlarged gape, facilitated by a low coronoid process and enlarged posterior mandibular tubercles (bony projections originating from the lower jaw), which provided an insertion site for sternum-to-mandible jaw abduction muscles, allowing for a more forceful opening of the jaw (Foss, 2001). All together, such traits suggest a massive and incredibly fearsome bite, perhaps the most formidable of any animal in its environment. Of course, none of such traits are especially indicative of a predatory lifestyle. Indeed, many modern non-predatory ungulates, like hippos, pigs and peccaries, also possess large, formidable skulls and jaws. However, in peeling back the layers, it is found there was more to the skull of Archaeotherium that lies in store. Indeed, when inspecting the animal closely, a unique mosaic of features is revealed; traits that make it out to be much more lethal than the average artiodactyl. On one hand, Archaeotherium possessed many traits similar to those of herbivores animals, as is expected of ungulates. For instance, its jaw musculature that allowed the lower jaw of Archaeotherium a full side-to-side chewing motion as in herbivores (whereas most carnivores can only move their lower jaw up and down)(Effinger, 1998). On the other hand, Archaeotherium wielded many other traits far more lethal in their morphology, less akin to a herbivore and far more akin to a bonafide predator. For instance, the aforementioned enlarged gape of Archaeotherium is a bizarre trait on a supposed herbivore, as such animals do not need large gapes to eat vegetation and thus have smaller, more restricted gapes. Conversely, many predatory lineages have comparatively large gapes, as larger gapes allow for the the jaws to grab on to more effectively larger objects, namely large prey animals (Joeckel, 1990). Such a juxtaposition, however, is most evident when discussing the real killing instruments of Archaeotherium — the teeth. More so than any facet of this animal, the teeth of Archaeotherium are the real stars of the show, showing both how alike it was compared to its herbivores counterparts and more importantly, how it couldn’t be more different. For instance, the molars of Archaeotherium were quite similar to modern herbivores ungulates, in that they were robust, bunodont, and were designed for crushing and grinding, similar in form and function to modern ungulates like peccaries (Joeckel, 1990). However, while the molars give the impression that Archaeotherium was a herbivore, the other teeth tell a very different story. The incisors, for example, were enlarged, sharpened, and fully interlocked (as opposed to the flat-topped incisors seen in herbivores ungulates), creating an incisor array that was seemingly ill-suited for cropping vegetation and much more adept at for gripping, puncturing and cutting (Joeckel, 1990). Even more formidable were the canines. Like the modern pigs from which entelodonts derived their nicknames, the canines of Archaeotherium were sharp and enlarged to form prominent tusk-like teeth, but unlike pigs, they were rounded in cross-section (similar to modern carnivores like big cats, indicating more durable canines that can absorb and resist torsional forces, such as those from struggling prey) and were serrated to form a distinct cutting edge (Effinger, 1998; Joeckel, 1990; Ruff & Van Valkenburgh, 1987). These canines, along with the incisors, interlock to stabilize the jaws while biting and dismantling in a carnivore-like fashion. More strikingly, the canines also seem to act as “occlusal guides,” wherein the canines help align the movement and position of the rear teeth as they come together, allowing for a more efficient shearing action by the rear teeth. This function is seen most prevalently modern carnivores mammals, and is evidenced by the canine tooth-wear, which is also analogous to modern predators like bears and canids (Joeckel, 1990). Indeed, going off such teeth alone, it is clear that Archaeotherium is far more predatory than expected of an ungulate. However, the real stars of the show, the teeth that truly betray the predatory nature of these ungulates, are the premolars. Perhaps the most carnivore-like teeth in the entelodont’s entire tooth row, the premolars of Archaeotherium, particularly the anterior premolars, are laterally compressed, somewhat conical in shape, and are weakly serrated to bear a cutting edge, giving them a somewhat carnivorous form and function of shearing and slicing (Effinger, 1998). Most strikingly of all, the premolars of Archaeotherium bear unique features similar not to modern herbivores, but to durophagous carnivores like hyenas, particularly apical wear patterns, highly thickened enamel, “zigzag-shaped” enamel prism layers (Hunter-Schraeger bands) on the premolars which is also seen in osteophagous animals like hyenas, and an interlocking premolar interface wherein linear objects (such as bones) inserted into jaws from the side would be pinned between the premolars and crushed (Foss, 2001). Taken together, these features do not suggest a diet of grass or vegetation like other ungulates. Rather, they suggest a far more violent diet, one including flesh as well as hard, durable foods, particularly bone. All in all, the evidence is clear. Archaeotherium and other entelodonts, unlike the rest of their artiodactyl kin, were not the passive herbivores as we envision ungulates today. Rather, they were willing, unrepentant meat-eaters that had a taste for flesh as well as foliage. Of course, even with such lines of evidence, its hard to conclude that Archaeotherium was a true predator. After all, its wide gape and durophagous teeth could have just as easily been used for scavenging or even to eat tough plant matter such as seeds or nuts, as in peccaries and pigs, which themselves share many of the same adaptations as Archaeotherium, include the more carnivorous ones (e.g. the wide gape, using the canines as an occlusal guide, etc.). How exactly do we know that these things were veritable predators and not pretenders to the title. To this end, there is yet one last piece of evidence, one that puts on full display the predatory prowess of Archaeotherium —evidence of a kill itself. Found within oligocene-aged sediment in what is now Wyoming, a collection of various fossil remains was found, each belonging to the ancient sheep-sized camel Poebrotherium, with many of the skeletal remains being disarticulated and even missing whole hindlimbs or even entire rear halves of their body. Tellingly, many of the remains bear extensive bite marks and puncture wounds across their surface. Upon close examination, the spacing and size of the punctures leave only one culprit: Archaeotherium. Of course, such an event could still have been scavenging; the entelodonts were consuming the remains of already dead, decomposed camels, explaining the bite marks. What was far more telling, however, was where the bite marks were found. In addition bite marks being found on the torso and lumbar regions of the camels, various puncture wounds were found on the skull and neck, which were otherwise uneaten. Scavengers rarely feast on the head to begin with; there is very little worthwhile meat on it besides the brain, cheek-muscles and eyes, and even if they did feed on the skull and neck, they would still eat it wholesale, not merely bite it and then leave it otherwise untouched. Indeed, it was clear that this was no mere scavenging event. Rather than merely consuming these camels, Archaeotherium was actively preying upon and killing them, dispatching them via a crushing bite to the skull or neck before dismembering and even bisecting the hapless camels with their powerful jaws to preferentially feast on their hindquarters (likely by swallowing the hindquarters whole, as the pelvis of Poebrotherium was coincidentally the perfect width for Archaeotherium to devour whole), eventually discarding the leftovers in meat caches for later consumption (Sundell, 1999). With this finding, such a feat of brutality leaves no doubt in ones mind as to what the true nature of Archaeotherium was. This was no herbivore, nor was it a simple scavenger. This was an active, rapacious predator, the most powerful in its entire ecosystem. Indeed, with such brutal evidence of predation frozen in time, combined with various dental, cranial, and post cranial adaptations of this formidable animal, it’s possible to paint a picture of how this formidable creature lived. Though an omnivore by trade, willing and able to feast on plant matter such as grass, roots and tubers, Archaeotherium was also a wanton predator that took just about any prey it wanted. Upon detecting its prey, it approached its vicim from ambush before launching itself at blazing speed. From there, its cursorial, hoofed legs, used by other ungulates for escape predation, were here employed to capture prey, carrying it at great speeds as it caught up to its quarry. Having closed the distance with its target, it was then that the entelodont brought its jaws to bear, grabbing hold of the victim with powerful jaws and gripping teeth to bring it to a screeching halt. If the victim is lucky, Archaeotherium will then kill it quickly with a crushing bite to the skull or neck, puncturing the brain or spinal cord and killing its target instantly. If not, the victim is eaten alive, torn apart while it’s still kicking, as modern boars will do today. In any case, incapacitated prey are subsequently dismantled, with the entelodont using its entire head and heavily-muscled necks to bite into and pull apart its victim in devastating “puncture-and pull’ bites (Foss, 2001). Prey would then finally be consumed starting at the hindquarters, with not even the bones of its prey being spared. Such brutality, though far from clean, drove home a singular truth: that during this time, ungulates were not just prey, that they were not the mere “predator-fodder” we know them as today. rather, they themselves were the predators themselves, dominating as superb hunters within their domain and even suppressing clades we know as predators today, least of all the carnivorans. Indeed, during this point in time, the age of the carnivorous ungulates had hit their stride, and more specifically, the age of entelodonts had begun. Of course, more so than any other ettelodont, Archaeotherium took to this new age with gusto. Archaeotherium lived from 35-28 million years ago during the late Eocene and early Oligocene in a locality known today as the White River Badlands, a fossil locality nestled along the Great Plains and Rocky Mountains. Though a chalky, barren landscape today, during the time of Archaeotherium, the White River Badlands was a swamp-like floodplain crisscrossed with rivers and interspersed with by a mosaic of forests concentrated around waterways, open woodlands and open plains. As with most ecosystems with such a lush disposition, this locale teemed with life, with ancient hornless rhinos, small horse-like hyracodonts and early camels roaming the open habitats while giant brontotheres, small early horses and strange, sheep-like ungulates called merycoidodonts (also known as “oreodonts”) dwelled within the dense forests. Within this locale, Archaeotherium stalked the open woodlands and riparian forests of its domain. Here, it acted as a dominant predator and scavenger across is territory, filling a niche similar to modern grizzly bears but far more predatory. Among its preferred food items would be plant matter such as roots, foliage and nuts, but also meat in the form of carrion or freshly caught prey. In this respect, smaller ungulates such as the fleet-footed camel Poebrotherium, a known prey item of Archaeotherium, would have made a for choice prey, as its small size would make it easy for Archaeotherium to dispatch with its powerful jaws, while the entelodonts swift legs gave it the speed necessary to keep pace with its agile prey. However, the entelodont didn’t have such a feast all to itself. Just as the badlands teemed with herbivores, so too did it teem with rival predators. Among their ranks included fearsome predators such as Hyaenodon, a powerful, vaguely dog-like predator up to the size of wolves (as in H. horridus) or even lions (as in the Eocene-aged H. megaloides, which was replaced by H. horridus during the Oligocene). Armed with a massive head, fierce jaws and a set of knife-like teeth that could cut down even large prey in seconds, these were some of the most formidable predators on the landscape. There were also the nimravids, cat-like carnivorans that bore saber-teeth to kill large prey in seconds, and included the likes of the lynx-sized Dinictis, the leopard-sized Hoplophoneus and even the jaguar-sized Eusmilus. Furthermore, there were amphicyonids, better known as the bear-dogs. Though known from much larger forms later on in their existence, during the late Eocene and Oligocene, they were much smaller and acted as the “canid-analogues” of the ecosystem, filling a role similar to wolves or coyotes. Last but not least, there were the bathornithid birds, huge cariamiform birds related to modern seriemas but much larger, which filled a niche similar to modern seriemas or secretary birds, albeit on a much larger scale. Given such competition, it would seem that Archaeotherium would have its hands full. However, things are not as they appear. For starters, habitat differences would mitigate high amounts of competition, as both Hyaenodon and the various nimravids occupy more specialized ecological roles (being a plains-specialist and forest-specialist, respectively) than did Archaeotherium, providing a buffer to stave off competition: More importantly, however, none of the aforementioned predators were simply big enough to take Archaeotherium on. During the roughly 7 million years existence of Archaeotherium, the only carnivore that matched it in size was H. megaloides, and even that would have an only applied to average A. mortoni individuals, not to the much larger, bison-sized “Megachoerus” individuals. The next largest predator at that point would be the jaguars-sized Eusmilus (specifically E. adelos) which would have only been a bit more than half the size of even an average A. mortoni. Besides that, virtually every other predator on the landscape was simply outclassed by the much larger entelodont in terms of size and brute strength. As such, within its domain, Archaeotherium had total, unquestioned authority, dominating the other predators in the landscape and likely stealing their kills as well. In fact, just about the only threat Archaeotherium had was other Archaeotherium, as fossil bite marks suggest that this animal regularly and fraglantly engaged in intraspecific combat, usually through face-biting and possibly even jaw-wrestling (Effinger, 1998; Tanke & Currie, 1998). Nevertheless, it was clear that Archaeotherium was the undisputed king of the badlands; in a landscape of hyaenodonts and carnivorans galore, it was a hoofed ungulate that reigned supreme. However, such a reign would not last. As the Eocene transitioned into the Eocene, the planet underwent an abrupt cooling and drying phase known as Eocene-Oligocene Transition or more simply the Grande Coupure. This change in climate would eliminate the sprawling wetlands and river systems that Archaeotherium had been depending on, gradually replacing it with drier and more open habitats. To its credit, Archaeotherium did manage to hang on, persisting well after the Grand-Coupure had taken place, but in the end the damage had been done; Archaeotherium was a dead-man-walking. Eventually, by around 28 million years ago, Archaeotherium would go extinct, perishing due to this change in global climate (Gillham, 2019). Entelodonts as a whole would persist into the Miocene, producing some of their largest forms ever known in the form of the bison-sized Daeodon (which was itself even more carnivorous than Archaeotherium), however they too would meet the same fate as their earlier cousins. By around 15-20 million years ago, entelodonts as a whole would go extinct. However, while the entelodonts may have perished, this was not the end of carnivorous ungulates as a whole. Recall that the cetacodontamorphs, the lineage of artiodactyls that produced the entelodonts, left behind two living descendants. The first among them were the hippos, themselves fairly frequent herbivores. The second of such lineage, however, was a different story. Emerging out of South Asia, this lineage of piscivorous cetacodontamorphs, in a an attempt to further specialize for the fish-hunting lifestyle, began to delve further and further into the water, becoming more and more aquatic and the millennia passed by. At a certain point, these carnivorous artiodactlys had become something completely unrecognizable from their original hoofed forms. Their skin became hairless and their bodies became streamlined for life in water. Their hoofed limbs grew into giant flippers for steering in the water and their previously tiny tails became massive and sported giant tail flukes for aquatic propulsion. Their noses even moved to the tip of their head, becoming a blowhole that would be signature to this clade as a whole. Indeed, this clade was none other than the modern whales, themselves derived, carnivorous ungulates that had specialized for a life in the water, and in doing so, became the some of the most dominant aquatic predators across the globe for millions of years. Indeed, though long gone, the legacy of the entelodonts and of predatory ungulates as a whole, a legacy Archaeotherium itself had helped foster, lives on in these paragons of predatory prowess, showing that the ungulates are more than just the mere “prey” that they are often made out to be. Moreover, given the success that carnivorous ungulates had enjoyed in the past and given how modern omnivorous ungulates like boar dabble in predation themselves, perhaps, in the distant future, this planet may see the rise of carnivorous ungulates once again, following in the footsteps left behind by Archaeotherium and the other predatory ungulates all those millions of years ago. |
2024.05.13 20:09 snowboardnirvana Google's Smartglasses Patent acquired from Canadian Company 'North' was published last week
2024.05.13 14:28 North-Accountant3231 Post surgery recovery and work life
Attaching my MRI here for reference. I have my appointment with my orthopedic doc shortly, but seems like i’m most likely headed towards a surgery, maybe a minimally invasive one with the camera? submitted by North-Accountant3231 to RotatorCuff [link] [comments] 27m, currently weighing 100kg. I lost around 15 kgs, and had built up a bit of strength/endurance over my last 6 months at the gym so a bit sad that will be put on halt for a while. I also have Cervical Radiculopathy due to a mild disc bulge at the C5-C6 level but my shoulder problems have been around since i was 21, due to a gym injury back in the day. Now i currently work as a software dev and I was wondering if I do indeed get surgery (which I think is the way forward but the doc will have the final word) what is my recovery looking like and how soon can I get back to work? My employers offer me the luxury of working from home, completely WFH too in case i’m unwell which I guess I qualify for now. I have a proper desk setup at home with an external monitor, keyboard, mouse, and a laptop stand for proper posture ever since my neck issues crept up. I am currently already going with physio for the neck, and have a few isometric exercises prescribed for the disc bulge. Given all this info, can anyone tell me what my road to recovery would look like and when I can get back into the gym? I’m not like a super athletic or bodybuilder, i just want to become healthy, have a certain level of cardiovascular fitness + some definition in the upper body since ive been chubby my entire life. Nothing too crazy but it sucks not being able to be active for a while. Also, more importantly than the gym, how soon can I get back to work? I would ideally not want to take more than a week or two off, has anyone had any experience typing with a sling on? Any help would be appreciated, thanks! |
2024.05.11 15:04 robeph [PC~Steam][2016-2021 (maybe)] Game where a group of characters explore and battle AI/Robots in 3/4 isometric in turn based combat
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2024.05.04 16:47 cloud_credits_throw Should I really be weight-bearing after 1 week post ACL + meniscus surgery?
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2024.04.29 10:26 VashxShanks [Steam Golden Week Big Sale 2024] List/Guide of Recommendations For Great JRPGs and Hidden Gems.
Game | Price/Link | Tags | Steam Deck |
---|---|---|---|
LIVE A LIVE | $24.99 | Turn-based/Different settings depending on the character chosen/HD-2D Graphics/min-tactical grid movement in battles/Remake of the SNES original/Amazing music | 🟦 Verified |
Persona 3 Reload | $48.99 | Turn-based/Modern Day setting/Highschool Life sim/Detective Mystery/Dating Sim/Social Links system/Great Soundtrack | 🟦 Verified |
Persona 4 Golden | $11.99 | Turn-based/Modern Day setting/Highschool Life sim/Detective Mystery/Dating Sim/Social Links system/Great Soundtrack | 🟦 Verified |
Persona 5 Royal | $23.99 | Turn-based/Modern Japan setting/Highschool life simulatoMystery/Monster collectoSocial link system/Great soundtrack | 🟦 Verified |
Soul Hackers 2 | $14.99 | Turn-based/Cyberpunk setting/Monster collectoFemale Protagonist | 🟧 Playable |
Yakuza: Like a Dragon | $11.99 | Turn-based/Contemporary Japan setting/Yakuza life/Open World/Heavy on Drama and Comedy/Heavy on Mini-games/Class Changing Mechanics | 🟦 Verified |
Like a Dragon: Ishin! | $23.99 | Action/Edo period Japan setting/Crime/Comedy heavy/Drama heavy/Mini-game Heavy/Beat'em up/Open World | 🟦 Verified |
Super Robot Wars 30 | $19.79 | Tactical turn-based/Sci-fi setting/Mecha/Anime CrossoveMission based progression/Huge list of Playable characters/VN style | 🟦 Verified |
Valkyria Chronicles | $4.99 | Tactical Turn-based/World War Militar setting/Tactical mixed with real-time elements/Sketch or "Canvas" art style/Build your Army with character customization/Mission based story progression/Army management | 🟦 Verified |
Valkyria Chronicles 4 Complete Edition | $9.99 | Tactical Turn-based/Fantasy Sci-fi World War setting/Tactical mixed with real-time elements/Sketch or "Canvas" art style/Build your Army with character customization/Mission based story progression/Army management | 🟦 Verified |
Labyrinth of Refrain: Coven of Dusk | $12.49 | Turn-based/Dark Fantasy setting/Class customization system/Tiered loot/Dungeon Crawler | 🟧 Playable |
Labyrinth of Galleria: The Moon Society | $39.99 | Turn-based/Dark Fantasy setting/Class customization system/Tiered loot/Dungeon Crawler | 🟦 Verified |
Langrisser I & II | $14.99 | Tactical Turn-based/Fantasy setting/Remake of 2 games in 1/Fire Emblem-like/Multiple story routes and endings | 🟦 Verified |
ŌKAMI HD | $9.99 | Action/Japanese Mythology Fantasy setting/Female Protagonist/Beautiful watercolor animations and art style | 🟦 Verified |
Monster Rancher 1 & 2 DX | $20.99 | Active time battle/Fantasy setting/Monster raising simulatoTournaments heavy/Monster collector | 🟦 Verified |
CRYSTAR | $17.99 | Action/Dark Fantasy setting/Female Protagonist/Hack & Slash/Deals with self-harm and dark subjects | 🟦 Verified |
The Caligula Effect: Overdose | $12.49 | Turn-based unique combat/School Life setting/Persona-like/Great Soundtrack/Female and Male Protagonist choice | -- |
Conception PLUS: Maidens of the Twelve Stars | $11.99 | Turn-based/Fantasy setting/Dating-sim/Fan-service/Harem/Dungeon Crawler | -- |
Conception II: Children of the Seven Stars | $3.99 | Turn-based/Fantasy setting/Dating-sim/Fan-service/Harem/Dungeon Crawler | -- |
The Alliance Alive HD Remastered | $9.99 | Turn-based/Fantasy setting/Expansive Skill Tree/Character customization | 🟦 Verified |
Zanki Zero: Last Beginning | $11.99 | Action/Post-apocalyptic setting/Base Building/Psychological HorroDating sim/Resource gathering & Survival/Crafting | 🟦 Verified |
MONARK | $38.99 | Tactical turn-based/Modern world setting/Dark Fantasy/Highschool setting/Psychological | -- |
Relayer Advanced | $18.99 | Tactical turn-based/Sci-fi setting/Space travel/Piloted Mecha/Female Protagonist/Anime art style | 🟧 Playable |
GOD WARS The Complete Legend | $5.99 | Tactical turn-based/Japanese Myth Fantasy setting/Class customization system | 🟦 Verified |
Cladun Returns Bundle (2 games): This Is Sengoku! & Cladun X2 | $4.96 | Action/Fantasy setting/Character creatoDungeon CrawlePixel graphics/Class system | 🟧 Playable |
Dust: An Elysian Tail | $4.94 | Action/Fantasy setting/Metriodvania/PlatformeMetroidvania | 🟦 Verified |
VALKYRIE ELYSIUM | $23.99 | Action/Fantasy setting/Female Protagonist/Multiple Endings | -- |
Hero Must Die. Again | $9.99 | Turn-based/Medieval Fantasy setting/choices MatteMany Different ending with each new playthrough/Short (4 to 5 hours) for each playthrough/Multiple playthroughs to get the true ending | 🟦 Verified |
The DioField Chronicle | $23.99 | Real Time Strategy/Medieval fantasy setting/Military & WaClass mechanics | 🟦 Verified |
SD GUNDAM G Generation Cross Rays | $9.99 | Tactical turn-based/Sci-fi setting/Mecha/Gundam Anime/Manga CrossoveMission based progression/Huge amount of playable Units & Pilots/Heavy & detailed customization options/Beautifully Animated | -- |
SD GUNDAM BATTLE ALLIANCE | $19.79 | Action/Sci-fi setting/Anime Adaptation/Hack and Slash/Online Co-Op | -- |
DRAGON QUEST® XI S: Echoes of an Elusive Age™ - Definitive Edition | $23.99 | Turn-based/Fantasy setting/Great soundtrack/Full Pixel graphics option/Character skill tree/Gathering resources and crafting | 🟧 Playable |
DRAGON QUEST BUILDERS™ 2 | $29.99 | Action/Fantasy setting/Minecraft-like | -- |
DRAGON QUEST TREASURES | $29.99 | Action/Medieval Fantasy setting/Monster CollectoTreasure hunteOpen Zones/Exploration | 🟦 Verified |
Secret of Mana | $19.99 | Action/Fantasy setting/3D Remake/Local Co-Op | -- |
Trials of Mana | $24.99 | Action/Fantasy setting/Hack & Slash/Choose 3 out of 6 main characters/Class customization system/Expansive Skill Tree | 🟦 Verified |
Legend of Mana Remaster | $14.99 | Action/Fantasy setting/Beat'em up/World Building Mechanic/Open World/Beautifully Hand Drawn/Great Music/Resource gathering & Crafting | 🟦 Verified |
AKIBA'S TRIP: Hellbound & Debriefed | $7.99 | Action/Modern Akihabara setting/Vampires/Beat'em up/Open-world/Comedy | 🟦 Verified |
AKIBA'S TRIP: Undead & Undressed (2nd game in the series) | $7.99 | Action/Modern Akihabara setting/Vampires/Beat'em up/Open-world/Comedy | -- |
NieR:Automata | $15.99 | Action/Hack & Slash/Female Protagonist/Post-apocalyptic setting/Open World/Dark Fantasy | 🟧 Playable |
NieR Replicant ver.1.22474487139... | $23.99 | Action/Post-apocalyptic setting/Hack & Slash/Bullet Hell/Dark Fantasy/Dark HumoFan-service/LGBTQ+ | 🟧 Playable |
Tactics Ogre: Reborn | $24.99 | Tactical turn-based/Medieval fantasy/Choices MatteMultiple story routes/Class system/Dark/Politics/Remake of the SNES original | 🟦 Verified |
Octopath Traveler | $29.99 | Turn-based/Fantasy setting/Multiple Protagonists to choose as your main charactePixel Graphics/2.5D | 🟧 Playable |
Octopath Traveler II | $35.99 | Turn-based/Fantasy setting/Multiple Protagonists to choose as your main charactePixel Graphics/2.5D | 🟦 Verified |
Bravely Default II | $29.99 | Turn-based/Fantasy setting/Class changing mechanics | 🟧 Playable |
TRIANGLE STRATEGY™ | $29.99 | Tactical Turn-based/Medieval Fantasy setting/HD2D/Choices MatteClass promotion mechanic/War & Politics/Multiple Story routes | 🟦 Verified |
HARVESTELLA | $29.99 | Action/Fantasy setting/Farming SimulatoClass changing system/Resource gathering and Crafting | 🟧 Playable |
Fallen Legion Revenants | $25.99 | Active Timed Battle/Fantasy setting/Valkyrie Profile combat-like/Choices Matter | -- |
Dungeon Encounters | $11.99 | Turn-based/Fantasy setting/Dungeon CrawleCharacter Customization/Very light on Story | 🟦 Verified |
Tales of Arise: Deluxe Edition | $19.99 | Action/Fantasy setting/Anime style/Revolution Story/Fantasy setting/anime trope heavy/Social links mechanic between party members | 🟦 Verified |
Chrono Trigger | $7.49 | Turn-based/Pixel Graphics/Time-Travel/Fantasy Adventure/Great Soundtrack/All time Classic | 🟧Playable |
NEO: The World Ends with You | $23.99 | Action/Modern Tokyo setting/Dark Fantasy/Death Game/Read people's minds/Psychic powers | 🟧Playable |
STAR OCEAN™ - THE LAST HOPE -™ 4K & Full HD Remaster | $6.29 | Action/Sci-fi setting/Space travel/Social link mechanic/Crafting and Resource gathering | -- |
Star Ocean The Divine Force | $29.99 | Action/Sci-fi mixed with Fantasy setting/Space travel/Resource gathering & Crafting/Social link mechanic between party members | -- |
Soul Nomad & the World Eaters | $9.99 | Tactical Turn-based/Fantasy setting/Deep character customization/Comedy/Multiple routes & endings | 🟧Playable |
Phantom Brave PC | $3.99 | Tactical Turn-based/Fantasy setting/Deep character customization/Comedy | 🟧Playable |
Makai Kingdom: Reclaimed and Rebound | $12.99 | Tactical Turn-based/Fantasy setting/Deep character customization/Comedy | -- |
ZHP: Unlosing Ranger vs. Darkdeath Evilman | $12.99 | Tactical Turn-based/Sci-fi setting/Deep character customization/Comedy/Roguelike | 🟦 Verified |
La Pucelle: Ragnarok | $12.99 | Tactical turn-based/Fantasy setting/Female Protagonist/Comedy | 🟦 Verified |
Rhapsody: A Musical Adventure | $12.99 | Tactical turn-based/Fantasy setting/Female Protagonist/Comedy/Cute/Musical | 🟦 Verified |
Rhapsody II: Ballad of the Little Princess | $19.99 | Tactical turn-based/Fantasy setting/Female Protagonist/Comedy/Cute/Musical | 🟦 Verified |
Rhapsody III: Memories of Marl Kingdom | $19.99 | Tactical turn-based/Fantasy setting/Female Protagonist/Comedy/Cute/Musical | 🟦 Verified |
Banner of the Maid | $1.69 | Tactical Turn-based/French War setting/Female Protagonist/Pixel Graphics | 🟦 Verified |
Persona 5 Tactica | $35.99 | Tactical turn-based/Isekai/Monster collector | 🟦 Verified |
Fate/Samurai Remnant | $40.19 | Action/Edo period Japan setting/Anime Visual Novel//Historic Figures | 🟦 Verified |
WitchSpring R | $27.29 | Turn-based/Medieval fantasy setting/Female Protagonist/Resource gathering & Crafting/Monster collection | 🟦 Verified |
Tales of Vesperia: Definitive Edition | $9.99 | Action/Fantasy setting/Anime style/Local Co-Op support in combat/Fantasy Adventure anime trope heavy | 🟦 Verified |
Tales of Arise | $19.99 | Action/Fantasy setting/Anime style/Revolution Story/Fantasy setting/anime trope heavy/Social links mechanic between party members | 🟦 Verified |
Blue Reflection | $14.99 | Turn-based/Japanese School Life setting/Female Protagonist/Magical Girls/Fan-service | -- |
Blue Reflection: Second Light | $23.99 | Turn-based/Japanese School Life setting/Female Protagonist/Magical Girls/Fan-service | 🟧 Playable |
Xanadu Next | $4.49 | Action/Fantasy setting/Isometric/Dungeon crawler | -- |
Loop8: Summer of Gods | $11.99 | Turn-based/Modern world setting/Social links/Dating sim/Choices matter | 🟦 Verified |
Regalia: Of Men and Monarchs | $3.99 | Tactical turn-based/Medieval Fantasy setting/Social link system/Politics/Comedy | 🟧 Playable |
Souls of Chronos | $9.59 | Action/Fantasy setting/Choices MatteMultiple Endings/Top-Down View/Chibi characteAnime art style | 🟦 Verified |
Demon Gaze Extra | $19.99 | Turn-based/First Person dungeon crawleDark Fantasy setting/Monster collection | 🟧 Playable |
Front Mission 1st: Remake | $17.49 | Tactical turn-based/Sci-fi setting/War & Politics/Mecha/Customization | 🟦Verified |
First off, I won't recommend big or famous games like Final Fantasy or Persona, this is more for lesser know, hidden gems, or just games where it's hard to find someone talking about them.
2024.04.28 11:23 MadCookie17 2 Tennis/2 Golfers elbow for years
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2024.04.24 19:04 HardFlaccidSyndrome0 Supplementary Tables
2024.04.23 21:49 Spirit198 Few Question On Overcoming Isometrics
2024.04.23 11:27 RomainMazB Teardrops of Mylen
Teardrops of MylenThe first roleplay forum really coupled with a browser based game designed like a TTRPG with DnD dice mechanism in mind.This simple description made you feel curious? Join our Discord. If not yet, continue reading! The same links will be available on the end of this speech! What makes this project uniqueThis web application is a roleplaying website with two main purposes:
The game is set up like a massively multiplayer TTRPG: a tiled map and various tokens. The main tokens are movement and action tokens, respectively 24 and 6 tokens reloaded gradually throughout the day which make the daily game time under 5 minutes (even under one if you want to be quick). An endless gameThe game basically never ends, there is no ultimate goal to reach that would end the game. Even "dying" from an enemy hit will just leave you on the floor and you can wait for an ally resurrection or choose to call your god reincarnate you to the map respawn area.Instead of a final goal, there is a competition system based on various statistics when you perform your actions. Many of these statistics are attached to a variety of pictograms that you collect with multiple rarity. Some pictograms are even distributed manually after winning a roleplaying contest or as a thank you for your participation in reporting bugs or suggestions. I need all of youI can talk for hours about the web development process or graphical AI generation, but roleplaying and community management is not my cup of tea.That is why I need a passionate community to actively participate in this project.I need players to make the forum and the game live, of course.But I also need people who want to actively participate behind the scenes like: improving the rules, organize roleplaying contests on the forum, getting new players feedback and answering their questions, help me to organize the community discord server, ... Want to participate?As of today, we are about to release an alpha test version this week to track down some bugs and get feedback about the game. To participate, you can join the Discord server and we'll give you the early access to the development environment so you can already create your character.Visual readers, here are some pics of the project.GameboardCharacter public profile page with pictograms to collect Character's actions history Ranking system Roleplay forum post |