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2024.05.14 18:08 Mophandel Archaeotherium, the King of the White River Badlands
Art by Bob Nicholls submitted by Mophandel to badassanimals [link] [comments] Nowadays, when we envision the words “prey,” among modern mammalian fauna, few taxa come to mind as quickly as the hoofed mammals, better known as the ungulates. Indeed, for the better part of their entire evolutionary history, the ungulates have become entirely indistinguishable from the term “prey.” Across their two major modern branches, the artiodactyls (the “even-toed ungulates,” such as bovids, pigs, deer, hippos and giraffes) and the perissodactyls (the “odd-toed ungulates,” including horses, rhinos and tapir), the ungulates too have created an empire spanning nearly every continent, establishing themselves as the the dominant herbivores throughout their entire range. However, as a price for such success, their lot as herbivores have forced them into an unenviable position: being the food for the predators. Indeed, throughout the diets of most modern predators, ungulates make up the majority, if not the entirety, of their diet, becoming their counterparts in this evolutionary dance of theirs. They have become the lamb to their wolf, the zebra to their lion, the stag to their tiger. If there is a predator in need of lunch, chances are that there is an ungulate there to provide it. Of course, such a dynamic is not necessarily a recent innovation. For the last 15-20 million years, across much of the world, both new and old, the ungulates have served as prey for these predators through it all. Over the course of whole epochs, these two groups have played into these roles for millions of years, coevolving with each other in an eons-long game of cat-and-mouse. The shoes they fill are not new, but have existed for ages, and within their niches they have cultivated their roles to perfection. Indeed, with such a tenured history, it seems hardly surprising the ungulates are wholly inseparable from the terms “prey,” itself. However, while this is the case now, as it has been for the last 15-20 million years, go back far enough, and we see that this dynamic is not as set in stone as we would think. Indeed, back during the Eocene and Oligocene, during the very earliest days of age of mammals, things were very different for the ungulates. While today they are considered little more than food for modern predators, during these olden days, the ungulates weren’t quite so benign. In fact, far from being fodder for top predators, the ungulates had turned the tables, instead becoming top predators themselves. Indeed, though nearly unheard of today, throughout much of the Eocene and Oligocene, carnivorous ungulates thrived in abundance, developing specializations for catching large prey and establishing themselves as top predators that competed alongside the more traditional carnivores, and even dominating them in some instances. Given such success, it’s no wonder that multiple such clades had arisen during this time. Such predators included the arctocyonids, a lineage of (ironically) hoof-less ungulates with large jaws and sharp teeth for capturing large prey. There were also the mesonychians, a lineage of dog-like ungulates with massive skulls and jaws that allowed them to reign as the top predator across much of the Eocene. However, among these various lineages, one stands stands out among the rest, by far. Arising during the Eocene, this lineage, though superficially resembling modern pigs, hailed from one an ancient lineage of artiodactyls far removed from swine or most other ungulates in general, with few close relatives alive today. Through perhaps not the most predatory of the bunch, it was among the most formidable, as their superficially pig-like appearance came with giant predatory jaws and teeth unlike anything from the modern era. And of course, as if all of that wasn’t enough, this lineage also went on to earn arguably one of the most badass nicknames of any lineage of mammals, period. These predators, of course, were the entelodonts, a.k.a the “hell-pigs.” More so than any other predatory ungulate lineage, these formidable ungulates were the ones to turn the current paradigm upside down, becoming some of the largest and most dominant carnivores in their landscape, even with (and often in spite of) the presence of more traditional predators. Through impressive size, fearsome teeth and sheer tenacity, these animals became the top dogs of their time, ruling as behemoth-kings of their Paleogene kingdoms, domineering all comers, and throughout the ranks, one entelodont in particular demonstrated such dominance the best. Though not the largest or most powerful of their kind, it is one of the most iconic, being among the most well-known members of its lineage to date. Moreover, this enteledont also has some of the most complete life histories ever seen out of this clade, with its brutality and predatory prowess being displayed in the fossil record in a way seen in no other member of its kind. More than anything else, however, it was this predator that best turned the notion of “ungulates being prey” on its head, living in an environment that bore some of the largest carnivoran hypercarnivores to date and still reigning as the undisputed top predator of its domain. This fearsome beast was none other than Archaeotherium, icon of the entelodonts, terror of the Oligocene American west and undisputed king of the White River badlands. The rise of Archaeotherium (and of entelodonts in general) is closely tied to the ascendancy of carnivorous ungulates as a whole, one of the earliest evolutionary success stories of the entire Cenozoic. Having become their own derived clade since the late Cretaceous, the ungulates were remarkably successful during the early Paleogene, as they were among the first mammalian clades to reach large sizes during those early days after the non-avian dinosaurs had gone extinct. As such, it was with incredible swiftness that, as the Paleogene progressed, the ungulates swooped upon the various niches left empty by the K-Pg mass extinction that killed the dinosaurs. This of course included the herbivorous niches we would know them for today, but this also included other, much more carnivore roles. Indeed, early on during the Paleogene, it was the ungulates that first seized the roles of large mammalian predators, becoming some the earliest large mammalian carnivores to ever live, well before even the carnivorans. Such predators included the arctocyonids, a lineage of vaguely dog-like, hoof-less ungulates with robust jaws and sharpened teeth that acted as some of earliest large carnivores of the Paleocene, with genera such as Arctocyon mumak getting up to the size of big cats. Even more prolific were the mesonychids. More so than what pretty much any other lineage of predator, it was the mesonychids that would stand out as the earliest dominant predators of the early Cenozoic. Growing up to the size of bears and with enormous, bone-crushing jaws, the mesonychids were among the most powerful and successful predators on the market at that time, with a near-global range and being capable of subjugating just about any other predator in their environments. Indeed, they, along with other carnivorous ungulates (as well as ungulates in general), were experiencing a golden age during this time, easily being the most prolific predators of the age. Given such prevalence, it should be no surprise that there would be yet another lineage of predatory ungulates would throw their hat into the ring, and by early Eocene, that contender would none other than the entelodonts. The very first entelodonts had arisen from artiodactyl ancestors during the Eocene epoch, at a time when artiodactyls were far more diverse and bizarre than they are now. Through today known from their modern herbivorous representatives such as bovines, deer, and antelope, during the Paleocene and Eocene, the artiodacyls, as with most ungulates of that time, were stronger and far more predaceous, particularly when it came to one such clade of artiodactyls, the cetacodontamorphs. Only known today from hippos and another group of artiodactyls (one which will become relevant later), the cetacodantomorphs emerged out of Asia around 55 million years ago, at around the same time that artiodactyls themselves had made their debut. These animals included the first truly predatory artiodactyls, with many of them possessing large skulls with powerful jaws and sharp, predatory teeth. Among their ranks included animals as puny as Indohyus, a piscivorous artiodactyl the size of a cat, to as formidable as Andrewsarchus, a giant, bison-sized predator often touted as one of the largest predatory mammals to ever live. Given such a predatory disposition, it wouldn’t be long until this clade produced a lineage of truly diverse, truly successful predators, and by around 40 million years ago, that is exactly what they did, as it was at that time that the entelodonts themselves first emerged. From their Asian homeland, the entelodonts spread across the world, spreading through not only most of Eurasia but also colonizing North America as well, with genera such as Brachyhyops being found across both continents. Here, in this North American frontier, the entelodonts began to diversify further, turning into their most successful and formidable forms yet, and it was around the late Eocene and early Oligocene that Archaeotherium itself had entered the scene. Just from a passing glance at Archaeotherium, it is clear how exactly it (as well as the other entelodonts) earned the nickname of “hell-pigs.” It was a bruiser for starters; its body bore a robust, pig-like physique, with prominent neural spines and their associated musculature forming a hump around the shoulder region, similar to the hump of a bison. With such a bulky physique came with it impressive size; the average A. mortoni had a head-body length of roughly 1.6-2.0 m (5.3-6.6 ft), a shoulder height of 1.2 m (4 ft) and a body mass of around 180 kg (396 lb) in weight (Boardman & Secord, 2013; Joeckel, 1990). At such sizes, an adult Archaeotherium the size of a large male black bear. However, they had the potential to get even bigger. While most Archaeotherium specimens were around the size described above, a select few specimens, labeled under the synonymous genus “Megachoerus,” are found to be much larger, with skulls getting up to 66% longer than average A. mortoni specimens (Foss, 2001; Joeckel, 1990). At such sizes and using isometric scaling, such massive Archaeotherium specimens would attained body lengths over 2.5 m (8.2 ft) and would have reached weighs well over 500 kg (1100 lb), or as big as a mature male polar bear. Indeed, at such sizes, it is already abundantly evident that Archaeotherium is a force to be recorded with. However, there was more to these formidable animals than sheer size alone. Behind all that bulk was an astoundingly swift and graceful predator, especially in terms of locomotion. Indeed, the hoofed feet of Archaeotherium, along with other entelodonts, sported several adaptations that gave it incredible locomotive efficiency, essentially turning it into a speed demon of the badlands. Such adaptations include longer distal leg elements (e.g. the radius and tibia) than their proximal counterparts (e.g. the humerus and femur), fusion of the radius and ulna for increased running efficiency, the loss of the clavicle (collar-bone) to allow for greater leg length, the loss of the acromion to enhance leg movement along the fore-and-aft plane, the loss of digits to reduce the mass of the forelimb, the fusion of the ectocuneiform and the mesocuneiform wrist-bones, among many other such traits (Theodore, 1996) . Perhaps most significant of these adaptations is the evolution of the “double-pulley astragalus (ankle-bone),” a specialized modification of the ankle that, while restricting rotation and side-to-side movement at the ankle-joint, allows for greater rotation in the fore-and-aft direction, thus allowing for more more powerful propulsion from the limbs, faster extension and retraction of the limbs and overall greater locomotive efficiency (Foss, 2001). Of course, such a trait was not only found in entelodonts but in artiodactyls as a whole, likely being a response to predatory pressures from incumbent predatory clades arising at the same time as the artiodactyls (Foss, 2001). However, in the case of the entelodonts, such adaptations were not used for merely escaping predators. Rather, they were used to for another, much more lethal effect… Such notions are further reinforced by the entelodonts most formidable aspect, none either than their fearsome jaws, and in this respect, Archaeotherium excelled. Both for its size and in general, the head of Archaeotherium was massive, measuring 40-50 cm (1.3-1.6 ft) in length among average A. mortoni specimens, to up to 78 cm (~2.6 ft) in the larger “Megachoerus” specimens (Joeckel, 1990). Such massive skulls were supported and supplemented by equally massive neck muscles and ligaments, which attached to massive neural spines on the anterior thoracic vertebrae akin to a bisons hump as well as to the sternum, allowing Archaeotherium to keep its head aloft despite the skulls massive size (Effinger, 1998). Of course, with such a massive skull, it should come as no surprise that such skulls housed exceptionally formidable jaws as well, and indeed, the bite of Archaeotherium was an especially deadly one. Its zygomatic arches (cheek-bones) and its temporal fossa were enlarged and expanded, indicative of massive temporalis muscles that afforded Archaeotherium astoundingly powerful bites (Joeckel, 1990). This is further augmented by Archaeotherium’s massive jugal flanges (bony projections of the cheek), which supported powerful masseter muscles which enhanced chewing and mastication, as well as an enlarged postorbital bar that reinforced the skull against torsional stresses (Foss, 2001). Last but not least, powerful jaws are supplemented by an enlarged gape, facilitated by a low coronoid process and enlarged posterior mandibular tubercles (bony projections originating from the lower jaw), which provided an insertion site for sternum-to-mandible jaw abduction muscles, allowing for a more forceful opening of the jaw (Foss, 2001). All together, such traits suggest a massive and incredibly fearsome bite, perhaps the most formidable of any animal in its environment. Of course, none of such traits are especially indicative of a predatory lifestyle. Indeed, many modern non-predatory ungulates, like hippos, pigs and peccaries, also possess large, formidable skulls and jaws. However, in peeling back the layers, it is found there was more to the skull of Archaeotherium that lies in store. Indeed, when inspecting the animal closely, a unique mosaic of features is revealed; traits that make it out to be much more lethal than the average artiodactyl. On one hand, Archaeotherium possessed many traits similar to those of herbivores animals, as is expected of ungulates. For instance, its jaw musculature that allowed the lower jaw of Archaeotherium a full side-to-side chewing motion as in herbivores (whereas most carnivores can only move their lower jaw up and down)(Effinger, 1998). On the other hand, Archaeotherium wielded many other traits far more lethal in their morphology, less akin to a herbivore and far more akin to a bonafide predator. For instance, the aforementioned enlarged gape of Archaeotherium is a bizarre trait on a supposed herbivore, as such animals do not need large gapes to eat vegetation and thus have smaller, more restricted gapes. Conversely, many predatory lineages have comparatively large gapes, as larger gapes allow for the the jaws to grab on to more effectively larger objects, namely large prey animals (Joeckel, 1990). Such a juxtaposition, however, is most evident when discussing the real killing instruments of Archaeotherium — the teeth. More so than any facet of this animal, the teeth of Archaeotherium are the real stars of the show, showing both how alike it was compared to its herbivores counterparts and more importantly, how it couldn’t be more different. For instance, the molars of Archaeotherium were quite similar to modern herbivores ungulates, in that they were robust, bunodont, and were designed for crushing and grinding, similar in form and function to modern ungulates like peccaries (Joeckel, 1990). However, while the molars give the impression that Archaeotherium was a herbivore, the other teeth tell a very different story. The incisors, for example, were enlarged, sharpened, and fully interlocked (as opposed to the flat-topped incisors seen in herbivores ungulates), creating an incisor array that was seemingly ill-suited for cropping vegetation and much more adept at for gripping, puncturing and cutting (Joeckel, 1990). Even more formidable were the canines. Like the modern pigs from which entelodonts derived their nicknames, the canines of Archaeotherium were sharp and enlarged to form prominent tusk-like teeth, but unlike pigs, they were rounded in cross-section (similar to modern carnivores like big cats, indicating more durable canines that can absorb and resist torsional forces, such as those from struggling prey) and were serrated to form a distinct cutting edge (Effinger, 1998; Joeckel, 1990; Ruff & Van Valkenburgh, 1987). These canines, along with the incisors, interlock to stabilize the jaws while biting and dismantling in a carnivore-like fashion. More strikingly, the canines also seem to act as “occlusal guides,” wherein the canines help align the movement and position of the rear teeth as they come together, allowing for a more efficient shearing action by the rear teeth. This function is seen most prevalently modern carnivorous mammals, and is evidenced by the canine tooth-wear, which is also analogous to modern predators like bears and canids (Joeckel, 1990). Indeed, going off such teeth alone, it is clear that Archaeotherium is far more predatory than expected of an ungulate. However, the real stars of the show, the teeth that truly betray the predatory nature of these ungulates, are the premolars. Perhaps the most carnivore-like teeth in the entelodont’s entire tooth row, the premolars of Archaeotherium, particularly the anterior premolars, are laterally compressed, somewhat conical in shape, and are weakly serrated to bear a cutting edge, giving them a somewhat carnivorous form and function of shearing and slicing (Effinger, 1998). Most strikingly of all, the premolars of Archaeotherium bear unique features similar not to modern herbivores, but to durophagous carnivores like hyenas, particularly apical wear patterns, highly thickened enamel, “zigzag-shaped” enamel prism layers (Hunter-Schraeger bands) on the premolars which is also seen in osteophagous animals like hyenas, and an interlocking premolar interface wherein linear objects (such as bones) inserted into jaws from the side would be pinned between the premolars and crushed (Foss, 2001). Taken together, these features do not suggest a diet of grass or vegetation like other ungulates. Rather, they suggest a far more violent diet, one including flesh as well as hard, durable foods, particularly bone. All in all, the evidence is clear. Archaeotherium and other entelodonts, unlike the rest of their artiodactyl kin, were not the passive herbivores as we envision ungulates today. Rather, they were willing, unrepentant meat-eaters that had a taste for flesh as well as foliage. Of course, even with such lines of evidence, its hard to conclude that Archaeotherium was a true predator. After all, its wide gape and durophagous teeth could have just as easily been used for scavenging or even to eat tough plant matter such as seeds or nuts, as in peccaries and pigs, which themselves share many of the same adaptations as Archaeotherium, include the more carnivorous ones (e.g. the wide gape, using the canines as an occlusal guide, etc.). How exactly do we know that these things were veritable predators and not pretenders to the title. To this end, there is yet one last piece of evidence, one that puts on full display the predatory prowess of Archaeotherium —evidence of a kill itself. Found within oligocene-aged sediment in what is now Wyoming, a collection of various fossil remains was found, each belonging to the ancient sheep-sized camel Poebrotherium, with many of the skeletal remains being disarticulated and even missing whole hindlimbs or even entire rear halves of their body. Tellingly, many of the remains bear extensive bite marks and puncture wounds across their surface. Upon close examination, the spacing and size of the punctures leave only one culprit: Archaeotherium. Of course, such an event could still have been scavenging; the entelodonts were consuming the remains of already dead, decomposed camels, explaining the bite marks. What was far more telling, however, was where the bite marks were found. In addition bite marks being found on the torso and lumbar regions of the camels, various puncture wounds were found on the skull and neck, which were otherwise uneaten. Scavengers rarely feast on the head to begin with; there is very little worthwhile meat on it besides the brain, cheek-muscles and eyes, and even if they did feed on the skull and neck, they would still eat it wholesale, not merely bite it and then leave it otherwise untouched. Indeed, it was clear that this was no mere scavenging event. Rather than merely consuming these camels, Archaeotherium was actively preying upon and killing them, dispatching them via a crushing bite to the skull or neck before dismembering and even bisecting the hapless camels with their powerful jaws to preferentially feast on their hindquarters (likely by swallowing the hindquarters whole, as the pelvis of Poebrotherium was coincidentally the perfect width for Archaeotherium to devour whole), eventually discarding the leftovers in meat caches for later consumption (Sundell, 1999). With this finding, such a feat of brutality leaves no doubt in ones mind as to what the true nature of Archaeotherium was. This was no herbivore, nor was it a simple scavenger. This was an active, rapacious predator, the most powerful in its entire ecosystem. Indeed, with such brutal evidence of predation frozen in time, combined with various dental, cranial, and post cranial adaptations of this formidable animal, it’s possible to paint a picture of how this formidable creature lived. Though an omnivore by trade, willing and able to feast on plant matter such as grass, roots and tubers, Archaeotherium was also a wanton predator that took just about any prey it wanted. Upon detecting its prey, it approached its vicim from ambush before launching itself at blazing speed. From there, its cursorial, hoofed legs, used by other ungulates for escape predation, were here employed to capture prey, carrying it at great speeds as it caught up to its quarry. Having closed the distance with its target, it was then that the entelodont brought its jaws to bear, grabbing hold of the victim with powerful jaws and gripping teeth to bring it to a screeching halt. If the victim is lucky, Archaeotherium will then kill it quickly with a crushing bite to the skull or neck, puncturing the brain or spinal cord and killing its target instantly. If not, the victim is eaten alive, torn apart while it’s still kicking, as modern boars will do today. In any case, incapacitated prey are subsequently dismantled, with the entelodont using its entire head and heavily-muscled necks to bite into and pull apart its victim in devastating “puncture-and pull’ bites (Foss, 2001). Prey would then finally be consumed starting at the hindquarters, with not even the bones of its prey being spared. Such brutality, though far from clean, drove home a singular truth: that during this time, ungulates were not just prey, that they were not the mere “predator-fodder” we know them as today. rather, they themselves were the predators themselves, dominating as superb hunters within their domain and even suppressing clades we know as predators today, least of all the carnivorans. Indeed, during this point in time, the age of the carnivorous ungulates had hit their stride, and more specifically, the age of entelodonts had begun. Of course, more so than any other entelodont, Archaeotherium took to this new age with gusto. Archaeotherium lived from 35-28 million years ago during the late Eocene and early Oligocene in a locality known today as the White River Badlands, a fossil locality nestled along the Great Plains and Rocky Mountains. Though a chalky, barren landscape today, during the time of Archaeotherium, the White River Badlands was a swamp-like floodplain crisscrossed with rivers and interspersed with by a mosaic of forests concentrated around waterways, open woodlands and open plains. As with most ecosystems with such a lush disposition, this locale teemed with life, with ancient hornless rhinos, small horse-like hyracodonts and early camels roaming the open habitats while giant brontotheres, small early horses and strange, sheep-like ungulates called merycoidodonts (also known as “oreodonts”) dwelled within the dense forests. Within this locale, Archaeotherium stalked the open woodlands and riparian forests of its domain. Here, it acted as a dominant predator and scavenger across is territory, filling a niche similar to modern grizzly bears but far more predatory. Among its preferred food items would be plant matter such as roots, foliage and nuts, but also meat in the form of carrion or freshly caught prey. In this respect, smaller ungulates such as the fleet-footed camel Poebrotherium, a known prey item of Archaeotherium, would have made a for choice prey, as its small size would make it easy for Archaeotherium to dispatch with its powerful jaws, while the entelodonts swift legs gave it the speed necessary to keep pace with its agile prey. However, the entelodont didn’t have such a feast all to itself. Just as the badlands teemed with herbivores, so too did it teem with rival predators. Among their ranks included fearsome predators such as Hyaenodon, a powerful, vaguely dog-like predator up to the size of wolves (as in H. horridus) or even lions (as in the Eocene-aged H. megaloides, which was replaced by H. horridus during the Oligocene). Armed with a massive head, fierce jaws and a set of knife-like teeth that could cut down even large prey in seconds, these were some of the most formidable predators on the landscape. There were also the nimravids, cat-like carnivorans that bore saber-teeth to kill large prey in seconds, and included the likes of the lynx-sized Dinictis, the leopard-sized Hoplophoneus and even the jaguar-sized Eusmilus. Furthermore, there were amphicyonids, better known as the bear-dogs. Though known from much larger forms later on in their existence, during the late Eocene and Oligocene, they were much smaller and acted as the “canid-analogues” of the ecosystem, filling a role similar to wolves or coyotes. Last but not least, there were the bathornithid birds, huge cariamiform birds related to modern seriemas but much larger, which filled a niche similar to modern seriemas or secretary birds, albeit on a much larger scale. Given such competition, it would seem that Archaeotherium would have its hands full. However, things are not as they appear. For starters, habitat differences would mitigate high amounts of competition, as both Hyaenodon and the various nimravids occupy more specialized ecological roles (being a plains-specialist and forest-specialist, respectively) than did Archaeotherium, providing a buffer to stave off competition: More importantly, however, none of the aforementioned predators were simply big enough to take Archaeotherium on. During the roughly 7 million years existence of Archaeotherium, the only carnivore that matched it in size was H. megaloides, and even that would have an only applied to average A. mortoni individuals, not to the much larger, bison-sized “Megachoerus” individuals. The next largest predator at that point would be the jaguars-sized Eusmilus (specifically E. adelos) which would have only been a bit more than half the size of even an average A. mortoni. Besides that, virtually every other predator on the landscape was simply outclassed by the much larger entelodont in terms of size and brute strength. As such, within its domain, Archaeotherium had total, unquestioned authority, dominating the other predators in the landscape and likely stealing their kills as well. In fact, just about the only threat Archaeotherium had was other Archaeotherium, as fossil bite marks suggest that this animal regularly and fraglantly engaged in intraspecific combat, usually through face-biting and possibly even jaw-wrestling (Effinger, 1998; Tanke & Currie, 1998). Nevertheless, it was clear that Archaeotherium was the undisputed king of the badlands; in a landscape of hyaenodonts and carnivorans galore, it was a hoofed ungulate that reigned supreme. However, such a reign would not last. As the Eocene transitioned into the Eocene, the planet underwent an abrupt cooling and drying phase known as Eocene-Oligocene Transition or more simply the Grande Coupure. This change in climate would eliminate the sprawling wetlands and river systems that Archaeotherium had been depending on, gradually replacing it with drier and more open habitats. To its credit, Archaeotherium did manage to hang on, persisting well after the Grand-Coupure had taken place, but in the end the damage had been done; Archaeotherium was a dead-man-walking. Eventually, by around 28 million years ago, Archaeotherium would go extinct, perishing due to this change in global climate (Gillham, 2019). Entelodonts as a whole would persist into the Miocene, producing some of their largest forms ever known in the form of the bison-sized Daeodon (which was itself even more carnivorous than Archaeotherium), however they too would meet the same fate as their earlier cousins. By around 15-20 million years ago, entelodonts as a whole would go extinct. However, while the entelodonts may have perished, this was not the end of carnivorous ungulates as a whole. Recall that the cetacodontamorphs, the lineage of artiodactyls that produced the entelodonts, left behind two living descendants. The first among them were the hippos, themselves fairly frequent herbivores. The second of such lineage, however, was a different story. Emerging out of South Asia, this lineage of piscivorous cetacodontamorphs, in a an attempt to further specialize for the fish-hunting lifestyle, began to delve further and further into the water, becoming more and more aquatic and the millennia passed by. At a certain point, these carnivorous artiodactlys had become something completely unrecognizable from their original hoofed forms. Their skin became hairless and their bodies became streamlined for life in water. Their hoofed limbs grew into giant flippers for steering in the water and their previously tiny tails became massive and sported giant tail flukes for aquatic propulsion. Their noses even moved to the tip of their head, becoming a blowhole that would be signature to this clade as a whole. Indeed, this clade was none other than the modern whales, themselves derived, carnivorous ungulates that had specialized for a life in the water, and in doing so, became the some of the most dominant aquatic predators across the globe for millions of years. Indeed, though long gone, the legacy of the entelodonts and of predatory ungulates as a whole, a legacy Archaeotherium itself had helped foster, lives on in these paragons of predatory prowess, showing that the ungulates are more than just the mere “prey” that they are often made out to be. Moreover, given the success that carnivorous ungulates had enjoyed in the past and given how modern omnivorous ungulates like boar dabble in predation themselves, perhaps, in the distant future, this planet may see the rise of carnivorous ungulates once again, following in the footsteps left behind by Archaeotherium and the other predatory ungulates all those millions of years ago. |
2024.05.14 17:59 Then_Marionberry_259 MAY 14, 2024 MAG.TO MAG SILVER REPORTS FIRST QUARTER FINANCIAL RESULTS
https://preview.redd.it/w3wbxgjzze0d1.png?width=3500&format=png&auto=webp&s=c343619687a11525804d04f755c495d975b2050d submitted by Then_Marionberry_259 to Treaty_Creek [link] [comments] VANCOUVER, British Columbia, May 14, 2024 (GLOBE NEWSWIRE) -- MAG Silver Corp. (TSX / NYSE American: MAG) (“MAG”, or the “Company”) announces the Company’s unaudited consolidated financial results for the three months ended March 31, 2024 (“Q1 2024”). For details of the unaudited condensed interim consolidated financial statements of the Company for the three months ended March 31, 2024 (“Q1 2024 Financial Statements”) and management’s discussion and analysis for the three months ended March 31, 2024 (“Q1 2024 MD&A”), please see the Company’s filings on the System for Electronic Document Analysis and Retrieval Plus (“SEDAR+”) at ( www.sedarplus.ca ) or on the Electronic Data Gathering, Analysis, and Retrieval (“EDGAR”) at ( www.sec.gov ). All amounts herein are reported in $000s of United States dollars (“US$”) unless otherwise specified (C$ refers to Canadian dollars). KEY HIGHLIGHTS (on a 100% basis unless otherwise noted)
1 Adjusted EBITDA, total cash costs, cash cost per ounce, all-in sustaining costs, all-in sustaining cost per ounce and free cash flow are non-IFRS measures, please see below ‘ Non-IFRS Measures ’ section and section 12 of the Q1 2024 MD&A for a detailed reconciliation of these measures to the Q1 2024 Financial Statements. 2 Equivalent silver head grade and equivalent silver production have been calculated using the following price assumptions to translate gold, lead and zinc to “equivalent” silver head grade and “equivalent” silver production: $23/oz silver, $1,950/oz gold, $0.95/lb lead and $1.15/lb zinc. 3 Equivalent silver ounces sold have been calculated using realized price assumptions to translate gold, lead and zinc to “equivalent” silver ounces sold (metal quantity, multiplied by metal price, divided by silver price). Q1 2024 realized prices of $23.73/oz silver, $2,112.27/oz gold, $0.92/lb lead and $1.08/lb zinc. CORPORATE
4 Information contained in or otherwise accessible through the Company’s website, including the 2022 sustainability report and MAG Silver 2022 ESG Data Table, do not form part of this News Release and are not incorporated into this News Release by reference. EXPLORATION
https://preview.redd.it/qdr6asmzze0d1.png?width=720&format=png&auto=webp&s=aaedd0020755eaf3404201557cc9623acc995125 JUANICIPIO RESULTS All results of Juanicipio in this section are on a 100% basis, unless otherwise noted. Operating Performance The following table and subsequent discussion provide a summary of the operating performance of Juanicipio for the three months ended March 31, 2024 and 2023, unless otherwise noted. https://preview.redd.it/9txq8snzze0d1.png?width=720&format=png&auto=webp&s=0089cf57573e8834bdb641cf9e055619e6f6c5b2 (1) Equivalent silver head grades have been calculated using the following price assumptions to translate gold, lead and zinc to “equivalent” silver head grade: $23/oz silver, $1,950/oz gold, $0.95/lb lead and $1.15/lb zinc (Q1 2023: $21.85/oz silver, $1,775/oz gold, $0.915/lb lead and $1.30/lb zinc). (2) Equivalent silver payable ounces have been calculated using realized price assumptions to translate gold, lead and zinc to “equivalent” silver payable ounces (metal quantity, multiplied by metal price, divided by silver price). Q1 2024 realized prices of $23.73/oz silver, $2,112.27/oz gold, $0.92/lb lead and $1.08/lb zinc (Q1 2023 realized prices of $22.93/oz silver, $1,959.50/oz gold, $0.94/lb lead and $1.43/lb zinc). During the three months ended March 31, 2024 a total of 325,081 tonnes of ore were mined. This represents an increase of 45% over Q1 2023. Increases in mined tonnages at Juanicipio have been driven by the operational ramp up of the mine towards steady state targets. During the three months ended March 31, 2024 a total of 325,683 tonnes of ore were processed through the Juanicipio plant; no ore was processed at the nearby Fresnillo and Saucito processing plants (100% owned by Fresnillo). This represents an increase of 47% over Q1 2023. The increase in milled tonnage has been driven by the Juanicipio mill commissioning and operational ramp up to nameplate capacity over the course of 2023. The silver head grade and equivalent silver head grade for the ore processed in the three months ended March 31, 2024 was 476 g/t and 713 g/t, respectively (three months ended March 31, 2023: 363 g/t and 530 g/t, respectively). Head grades in Q1 2023 were lower as low-grade commissioning stockpiles were processed through the Juanicipio plant. Silver metallurgical recovery during Q1 2024 was 89.1% (Q1 2023: 87.0%) reflecting ongoing optimizations in the processing plant. The following table provides a summary of the total cash costs 5 and all-in sustaining costs 5 (“AISC”) of Juanicipio for the three months ended March 31, 2024, and 2023. https://preview.redd.it/riqz1yozze0d1.png?width=720&format=png&auto=webp&s=c5f2edde6bc789d2d379ca3a910beaa1cb262bde ________________________ 5 Total cash costs, cash cost per ounce, cash cost per equivalent ounce, all-in sustaining costs, all-in sustaining cost per ounce, and all-in sustaining cost per equivalent ounce are non-IFRS measures, please see the “ Non-IFRS Measures ” section below and section 12 of the Q1 2024 MD&A for a detailed reconciliation of these measures to the Q1 2024 Financial Statements. Equivalent silver ounces sold have been calculated using realized price assumptions to translate gold, lead and zinc to “equivalent” silver ounces sold (metal quantity, multiplied by metal price, divided by silver price). Q1 2024 realized prices of $23.73/oz silver, $2,112.27/oz gold, $0.92/lb lead and $1.08/lb zinc (Q1 2023: $22.93/oz silver, $1,959.50/oz gold, $0.94/lb lead and $1.43/lb zinc). Financial Results The following table presents excerpts of the financial results of Juanicipio for the three months ended March 31, 2024 and 2023. https://preview.redd.it/4zym5dqzze0d1.png?width=720&format=png&auto=webp&s=d25c91cabc7f6c6b35ff61acacc0160f3dfac2e7 Sales increased by $72,207 during the three months ended March 31, 2024, mainly due to 179% higher metal volumes and 2% higher realized metal prices. Offsetting higher sales was higher production cost ($9,409) which was driven by higher sales and operational ramp-up in mining and processing, including $3,545 in inventory movements, and higher depreciation ($14,083) as the Juanicipio mill achieved commercial production and commenced depreciating the processing facility and associated equipment in June 2023. Operating margin increased by 21% to 52%, mainly due to operational leverage and the lower reliance on the nearby Fresnillo and Saucito processing facilities. Other expenses increased by $2,159 mainly as a result of higher extraordinary mining and other duties ($872) in relation to higher precious metal revenues from the sale of concentrates and higher consulting and administrative expenses ($2,690) as an operator services agreement became effective upon initiation of commercial production (the “Operator Services Agreement”), offset by lower exchange losses and other costs ($1,566). Taxes increased by $20,980 impacted by higher taxable profits generated during Q1 2024, and non-cash deferred tax credits related to the commencement of use of plant and equipment in Q1 2023. Ore Processed at Juanicipio Plant (100% basis) https://preview.redd.it/koajybrzze0d1.png?width=720&format=png&auto=webp&s=caf1b70c53c47d371b9cebd743877fcd4ae59ba3 (1) The underground mine was considered readied for its intended use on January 1, 2022, whereas the Juanicipio processing facility started commissioning and ramp-up activities in January 2023, achieving commercial production status on June 1, 2023. (2) Includes toll milling costs from processing mineralized material at the Saucito and Fresnillo plants for Q1 2023. Sales and treatment charges are recorded on a provisional basis and are adjusted based on final assay and pricing adjustments in accordance with the offtake contracts. MAG FINANCIAL RESULTS – THREE MONTHS ENDED MARCH 31, 2024 As at March 31, 2024, MAG had working capital of $72,833 (December 31, 2023: $67,262) including cash of $74,683 (December 31, 2023: $68,707) and no long-term debt. As well, as at March 31, 2024, Juanicipio had working capital of $107,088 including cash of $30,991 (MAG’s attributable share is 44%). The Company’s net income for the three months ended March 31, 2024 amounted to $14,895 (March 31, 2023: $4,713) or $0.14/share (March 31, 2023: $0.05/share). MAG recorded its 44% income from equity accounted investment in Juanicipio of $19,244 (March 31, 2023: $7,919) which included MAG’s 44% share of net income from operations as well as loan interest earned on loans advanced to Juanicipio (see above for MAG’s share of income from its equity accounted investment in Juanicipio). https://preview.redd.it/5bneimszze0d1.png?width=720&format=png&auto=webp&s=323d988dbd92cf6ca4e815fff071c8ec7f45ee00 NON-IFRS MEASURES The following table provides a reconciliation of cash cost per silver ounce of Juanicipio to production cost of Juanicipio on a 100% basis (the nearest IFRS measure) as presented in the notes to the Q1 2024 Financial Statements. https://preview.redd.it/0jhh2stzze0d1.png?width=720&format=png&auto=webp&s=c97634763bd00967ddd878bc09b8e069687fae68 (1) As Q3 2023 represented the first full quarter of commercial production, information presented for total cash costs together with their associated per unit values are not directly comparable. (2) By-product revenues relates to the sale of other metals namely gold, lead, and zinc. (3) Equivalent silver payable ounces have been calculated using realized prices to translate gold, lead and zinc to “equivalent” silver payable ounces (metal quantity, multiplied by metal price, divided by silver price). Q1 2024 realized prices: $23.73/oz silver, $2,112.27/oz gold, $0.92/lb lead and $1.08/lb zinc (Q1 2023: $22.93/oz silver, $1,959.50/oz gold, $0.94/lb lead and $1.43/lb zinc). The following table provides a reconciliation of AISC of Juanicipio to production cost and various operating expenses of Juanicipio on a 100% basis (the nearest IFRS measure), as presented in the notes to the Q1 2024 Financial Statements. https://preview.redd.it/qkx2r5vzze0d1.png?width=720&format=png&auto=webp&s=0c9cb89471512649ac77b5cbe3177b0621d64835 (1) As Q3 2023 represented the first full quarter of commercial production, information presented for all-in sustaining costs and all-in sustaining margin together with their associated per unit values are not directly comparable. (2) Equivalent silver payable ounces have been calculated using realized prices to translate gold, lead and zinc to “equivalent” silver payable ounces (metal quantity, multiplied by metal price, divided by silver price). Q1 2024 realized prices: $23.73/oz silver, $2,112.27/oz gold, $0.92/lb lead and $1.08/lb zinc, (Q1 2023 realized prices: $22.93/oz silver, $1,959.50/oz gold, $0.94/lb lead and $1.43/lb zinc). For the three months ended March 31, 2024 the Company incurred corporate G&A expenses of $3,964 (three months ended March 31, 2023: $3,262), which exclude depreciation expense. The Company’s attributable silver ounces sold and equivalent silver ounces sold for the three months ended March 31, 2024 were 1,757,630 and 2,475,862 respectively (three months ended March 31, 2023: 880,429 and 1,230,412 respectively), resulting in additional all‐in sustaining cost for the Company of $2.26/oz and $1.60/oz respectively (three months ended March 31, 2023: $3.71/oz and $2.65/oz respectively), in addition to Juanicipio’s all-in-sustaining costs presented in the above table. The following table provides a reconciliation of EBITDA and Adjusted EBITDA attributable to the Company based on its economic interest in Juanicipio to net income (the nearest IFRS measure) of the Company per the Q1 2024 Financial Statements. All adjustments are shown net of estimated income tax. https://preview.redd.it/sx6jo7wzze0d1.png?width=720&format=png&auto=webp&s=1fea9df2b8195ce579b570935ab24b9939da5988 (1) As Q3 2023 represents the first full quarter of commercial production, information presented for EBITDA and Adjusted EBITDA is not directly comparable. The following table provides a reconciliation of free cash flow of Juanicipio to its cash flow from operating activities on a 100% basis (the nearest IFRS measure), as presented in the notes to the Q1 2024 Financial Statements. https://preview.redd.it/aqnp2dxzze0d1.png?width=720&format=png&auto=webp&s=56db2245eb87ec1f718edc1890b48b9b5d3ad762 (1) As Q3 2023 represents the first full quarter of commercial production, comparative information presented for free cash flow of Juanicipio is not directly comparable. Qualified Persons: All scientific or technical information in this press release including assay results referred to, and mineral resource estimates, if applicable, is based upon information prepared by or under the supervision of, or has been approved by Gary Methven, P.Eng., Vice President, Technical Services and Lyle Hansen, P.Geo, Geotechnical Director; both are “Qualified Persons” for purposes of National Instrument 43-101, Standards of Disclosure for Mineral Projects About MAG Silver Corp. MAG Silver Corp. is a growth-oriented Canadian exploration company focused on advancing high-grade, district scale precious metals projects in the Americas. MAG is emerging as a top-tier primary silver mining company through its (44%) joint venture interest in the 4,000 tonnes per day Juanicipio Mine, operated by Fresnillo plc (56%). The mine is located in the Fresnillo Silver Trend in Mexico, the world's premier silver mining camp, where in addition to underground mine production and processing of high-grade mineralised material, an expanded exploration program is in place targeting multiple highly prospective targets. MAG is also executing multi-phase exploration programs at the 100% earn-in Deer Trail Project in Utah and the 100% owned Larder Project, located in the historically prolific Abitibi region of Canada. Neither the Toronto Stock Exchange nor the NYSE American has reviewed or accepted responsibility for the accuracy or adequacy of this press release, which has been prepared by management. Certain information contained in this release, including any information relating to MAG’s future oriented financial information, are “forward-looking information” and “forward-looking statements” within the meaning of applicable Canadian and United States securities legislation (collectively herein referred as “forward-looking statements”), including the “safe harbour” provisions of provincial securities legislation, the U.S. Private Securities Litigation Reform Act of 1995, Section 21E of the U.S. Securities Exchange Act of 1934, as amended and Section 27A of the U.S. Securities Act. Such forward-looking statements include, but are not limited to:
Forward-looking statements are necessarily based upon estimates and assumptions, which are inherently subject to significant business, economic and competitive uncertainties and contingencies, many of which are beyond the Company’s control and many of which, regarding future business decisions, are subject to change. Assumptions underlying the Company’s expectations regarding forward-looking statements contained in this release include, among others: MAG’s ability to carry on its various exploration and development activities including project development timelines, the timely receipt of required approvals and permits, the price of the minerals produced, the costs of operating, exploration and development expenditures, the impact on operations of the Mexican tax and legal regimes, MAG’s ability to obtain adequate financing, outbreaks or threat of an outbreak of a virus or other contagions or epidemic disease will be adequately responded to locally, nationally, regionally and internationally. Although MAG believes the expectations expressed in such forward-looking statements are based on reasonable assumptions, such statements are not guarantees of future performance and actual results or developments may differ materially from those in the forward-looking statements. These forward-looking statements involve known and unknown risks, uncertainties and many factors could cause actual results, performance or achievements to be materially different from any future results, performance or achievements that may be expressed or implied by such forward-looking statements including amongst others: commodities prices; changes in expected mineral production performance; unexpected increases in capital costs or cost overruns; exploitation and exploration results; continued availability of capital and financing; general economic, market or business conditions; risks relating to the Company’s business operations; risks relating to the financing of the Company’s business operations; risks related to the Company’s ability to comply with restrictive covenants and maintain financial covenants pursuant to the terms of the Credit Facility; the expected use of the Credit Facility; risks relating to the development of Juanicipio and the minority interest investment in the same; risks relating to the Company’s property titles; risks related to receipt of required regulatory approvals; pandemic risks; supply chain constraints and general costs escalation in the current inflationary environment heightened by the invasion of Ukraine by Russia and the events relating to the Israel-Hamas war; risks relating to the Company’s financial and other instruments; operational risk; environmental risk; political risk; currency risk; market risk; capital cost inflation risk; risk relating to construction delays; the risk that data is incomplete or inaccurate; the risks relating to the limitations and assumptions within drilling, engineering and socio-economic studies relied upon in preparing economic assessments and estimates, including the 2017 PEA; as well as those risks more particularly described under the heading “Risk Factors” in the Company’s Annual Information Form dated March 27, 2023 available under the Company’s profile on SEDAR+ at www.sedarplus.ca . Should one or more of these risks or uncertainties materialize, or should underlying assumptions prove incorrect, actual results may vary materially from those described herein. This list is not exhaustive of the factors that may affect any of the Company’s forward-looking statements. The Company’s forward-looking statements are based on the beliefs, expectations and opinions of management on the date the statements are made and, other than as required by applicable securities laws, the Company does not assume any obligation to update forward-looking statements if circumstances or management’s beliefs, expectations or opinions should change. For the reasons set forth above, investors should not attribute undue certainty to or place undue reliance on forward-looking statements. Please Note: Investors are urged to consider closely the disclosures in MAG's annual and quarterly reports and other public filings, accessible through the Internet at www.sedarplus.ca and www.sec.gov LEI: 254900LGL904N7F3EL14 For further information on behalf of MAG Silver Corp. Contact Michael J. Curlook, Vice President, Investor Relations and Communications Phone: (604) 630-1399 Toll Free: (866) 630-1399 Email:info@magsilver.comhttps://preview.redd.it/d8wzg8yzze0d1.jpg?width=66&format=pjpg&auto=webp&s=a39ec6a39761447bbd4dbd77e92aef49e95e583e https://preview.redd.it/qr1wv0zzze0d1.png?width=4000&format=png&auto=webp&s=fadeb32c9245f8b70d903b1c70321decddf77985
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2024.05.14 17:48 water_elaborate 23M Bulgaria, looking for a weird one
2024.05.14 17:45 Horror-Cut-6375 My cat needs surgery is there any non profit that can help me? In RI
2024.05.14 17:18 MichaelPP3 Hustling Cargo (a Lucky Charm excerpt)
2024.05.14 17:10 No_Green_4369 AITA for questioning my husband's faithfulness?
2024.05.14 17:07 Ashamed-Egg-9180 My girlfriends (26F) sugar addiction is getting worse and I don't know what to do? (28M)
2024.05.14 16:58 TheRealStorey Just got a call about recommended maintenance.
2024.05.14 16:30 upbstock more Powell ------------
Fed Chair Powell said the economy has been performing very wellFed Chair Powell said the economy has been performing very well lately, in his comments from Amsterdam. He would not call the just-released PPI hot, and said it was rather quite mixed. He acknowledged that Q1 has been notable for the lack of inflation progress. It was not expected to be a smooth path lower, and he repeated that the Fed needs to be patient and let restrictive policy work. Monthly inflation rates are expected to move back down, but his confidence is not as high as it was after the downdraft in pressures last year. There are signs of cooling in the labor market, which is coming into better balance. The Fed does not know yet what it would do if inflation proved more persistent, but suggested it was likely that policy would be left where it is.
2024.05.14 16:25 Mophandel Archaeotherium, the King of the White River Badlands
Art by Bob Nicholls submitted by Mophandel to Naturewasmetal [link] [comments] Nowadays, when we envision the words “prey,” among modern mammalian fauna, few taxa come to mind as quickly as the hoofed mammals, better known as the ungulates. Indeed, for the better part of their entire evolutionary history, the ungulates have become entirely indistinguishable from the term “prey.” Across their two major modern branches, the artiodactyls (the “even-toed ungulates,” such as bovids, pigs, deer, hippos and giraffes) and the perissodactyls (the “odd-toed ungulates,” including horses, rhinos and tapir), the ungulates too have created an empire spanning nearly every continent, establishing themselves as the the dominant herbivores throughout their entire range. However, as a price for such success, their lot as herbivores have forced them into an unenviable position: being the food for the predators. Indeed, throughout the diets of most modern predators, ungulates make up the majority, if not the entirety, of their diet, becoming their counterparts in this evolutionary dance of theirs. They have become the lamb to their wolf, the zebra to their lion, the stag to their tiger. If there is a predator in need of lunch, chances are that there is an ungulate there to provide it. Of course, such a dynamic is not necessarily a recent innovation. For the last 15-20 million years, across much of the world, both new and old, the ungulates have served as prey for these predators through it all. Over the course of whole epochs, these two groups have played into these roles for millions of years, coevolving with each other in an eons-long game of cat-and-mouse. The shoes they fill are not new, but have existed for ages, and within their niches they have cultivated their roles to perfection. Indeed, with such a tenured history, it seems hardly surprising the ungulates are wholly inseparable from the terms “prey,” itself. However, while this is the case now, as it has been for the last 15-20 million years, go back far enough, and we see that this dynamic is not as set in stone as we would think. Indeed, back during the Eocene and Oligocene, during the very earliest days of age of mammals, things were very different for the ungulates. While today they are considered little more than food for modern predators, during these olden days, the ungulates weren’t quite so benign. In fact, far from being fodder for top predators, the ungulates had turned the tables, instead becoming top predators themselves. Indeed, though nearly unheard of today, throughout much of the Eocene and Oligocene, carnivorous ungulates thrived in abundance, developing specializations for catching large prey and establishing themselves as top predators that competed alongside the more traditional carnivores, and even dominating them in some instances. Given such success, it’s no wonder that multiple such clades had arisen during this time. Such predators included the arctocyonids, a lineage of (ironically) hoof-less ungulates with large jaws and sharp teeth for capturing large prey. There were also the mesonychians, a lineage of dog-like ungulates with massive skulls and jaws that allowed them to reign as the top predator across much of the Eocene. However, among these various lineages, one stands stands out among the rest, by far. Arising during the Eocene, this lineage, though superficially resembling modern pigs, hailed from one an ancient lineage of artiodactyls far removed from swine or most other ungulates in general, with few close relatives alive today. Through perhaps not the most predatory of the bunch, it was among the most formidable, as their superficially pig-like appearance came with giant predatory jaws and teeth unlike anything from the modern era. And of course, as if all of that wasn’t enough, this lineage also went on to earn arguably one of the most badass nicknames of any lineage of mammals, period. These predators, of course, were the entelodonts, a.k.a the “hell-pigs.” More so than any other predatory ungulate lineage, these formidable ungulates were the ones to turn the current paradigm upside down, becoming some of the largest and most dominant carnivores in their landscape, even with (and often in spite of) the presence of more traditional predators. Through impressive size, fearsome teeth and sheer tenacity, these animals became the top dogs of their time, ruling as behemoth-kings of their Paleogene kingdoms, domineering all comers, and throughout the ranks, one entelodont in particular demonstrated such dominance the best. Though not the largest or most powerful of their kind, it is one of the most iconic, being among the most well-known members of its lineage to date. Moreover, this enteledont also has some of the most complete life histories ever seen out of this clade, with its brutality and predatory prowess being displayed in the fossil record in a way seen in no other member of its kind. More than anything else, however, it was this predator that best turned the notion of “ungulates being prey” on its head, living in an environment that bore some of the largest carnivoran hypercarnivores to date and still reigning as the undisputed top predator of its domain. This fearsome beast was none other than Archaeotherium, icon of the entelodonts, terror of the Oligocene American west and undisputed king of the White River badlands. The rise of Archaeotherium (and of entelodonts in general) is closely tied to the ascendancy of carnivorous ungulates as a whole, one of the earliest evolutionary success stories of the entire Cenozoic. Having become their own derived clade since the late Cretaceous, the ungulates were remarkably successful during the early Paleogene, as they were among the first mammalian clades to reach large sizes during those early days after the non-avian dinosaurs had gone extinct. As such, it was with incredible swiftness that, as the Paleogene progressed, the ungulates swooped upon the various niches left empty by the K-Pg mass extinction that killed the dinosaurs. This of course included the herbivorous niches we would know them for today, but this also included other, much more carnivore roles. Indeed, early on during the Paleogene, it was the ungulates that first seized the roles of large mammalian predators, becoming some the earliest large mammalian carnivores to ever live, well before even the carnivorans. Such predators included the arctocyonids, a lineage of vaguely dog-like, hoof-less ungulates with robust jaws and sharpened teeth that acted as some of earliest large carnivores of the Paleocene, with genera such as Arctocyon mumak getting up to the size of big cats. Even more prolific were the mesonychids. More so than what pretty much any other lineage of predator, it was the mesonychids that would stand out as the earliest dominant predators of the early Cenozoic. Growing up to the size of bears and with enormous, bone-crushing jaws, the mesonychids were among the most powerful and successful predators on the market at that time, with a near-global range and being capable of subjugating just about any other predator in their environments. Indeed, they, along with other carnivorous ungulates (as well as ungulates in general), were experiencing a golden age during this time, easily being the most prolific predators of the age. Given such prevalence, it should be no surprise that there would be yet another lineage of predatory ungulates would throw their hat into the ring, and by early Eocene, that contender would none other than the entelodonts. The very first entelodonts had arisen from artiodactyl ancestors during the Eocene epoch, at a time when artiodactyls were far more diverse and bizarre than they are now. Through today known from their modern herbivorous representatives such as bovines, deer, and antelope, during the Paleocene and Eocene, the artiodacyls, as with most ungulates of that time, were stronger and far more predaceous, particularly when it came to one such clade of artiodactyls, the cetacodontamorphs. Only known today from hippos and another group of artiodactyls (one which will become relevant later), the cetacodantomorphs emerged out of Asia around 55 million years ago, at around the same time that artiodactyls themselves had made their debut. These animals included the first truly predatory artiodactyls, with many of them possessing large skulls with powerful jaws and sharp, predatory teeth. Among their ranks included animals as puny as Indohyus, a piscivorous artiodactyl the size of a cat, to as formidable as Andrewsarchus, a giant, bison-sized predator often touted as one of the largest predatory mammals to ever live. Given such a predatory disposition, it wouldn’t be long until this clade produced a lineage of truly diverse, truly successful predators, and by around 40 million years ago, that is exactly what they did, as it was at that time that the entelodonts themselves first emerged. From their Asian homeland, the entelodonts spread across the world, spreading through not only most of Eurasia but also colonizing North America as well, with genera such as Brachyhyops being found across both continents. Here, in this North American frontier, the entelodonts began to diversify further, turning into their most successful and formidable forms yet, and it was around the late Eocene and early Oligocene that Archaeotherium itself had entered the scene. Just from a passing glance at Archaeotherium, it is clear how exactly it (as well as the other entelodonts) earned the nickname of “hell-pigs.” It was a bruiser for starters; its body bore a robust, pig-like physique, with prominent neural spines and their associated musculature forming a hump around the shoulder region, similar to the hump of a bison. With such a bulky physique came with it impressive size; the average A. mortoni had a head-body length of roughly 1.6-2.0 m (5.3-6.6 ft), a shoulder height of 1.2 m (4 ft) and a body mass of around 180 kg (396 lb) in weight (Boardman & Secord, 2013; Joeckel, 1990). At such sizes, an adult Archaeotherium the size of a large male black bear. However, they had the potential to get even bigger. While most Archaeotherium specimens were around the size described above, a select few specimens, labeled under the synonymous genus “Megachoerus,” are found to be much larger, with skulls getting up to 66% longer than average A. mortoni specimens (Foss, 2001; Joeckel, 1990). At such sizes and using isometric scaling, such massive Archaeotherium specimens would attained body lengths over 2.5 m (8.2 ft) and would have reached weighs well over 500 kg (1100 lb), or as big as a mature male polar bear. Indeed, at such sizes, it is already abundantly evident that Archaeotherium is a force to be recorded with. However, there was more to these formidable animals than sheer size alone. Behind all that bulk was an astoundingly swift and graceful predator, especially in terms of locomotion. Indeed, the hoofed feet of Archaeotherium, along with other entelodonts, sported several adaptations that gave it incredible locomotive efficiency, essentially turning it into a speed demon of the badlands. Such adaptations include longer distal leg elements (e.g. the radius and tibia) than their proximal counterparts (e.g. the humerus and femur), fusion of the radius and ulna for increased running efficiency, the loss of the clavicle (collar-bone) to allow for greater leg length, the loss of the acromion to enhance leg movement along the fore-and-aft plane, the loss of digits to reduce the mass of the forelimb, the fusion of the ectocuneiform and the mesocuneiform wrist-bones, among many other such traits (Theodore, 1996) . Perhaps most significant of these adaptations is the evolution of the “double-pulley astragalus (ankle-bone),” a specialized modification of the ankle that, while restricting rotation and side-to-side movement at the ankle-joint, allows for greater rotation in the fore-and-aft direction, thus allowing for more more powerful propulsion from the limbs, faster extension and retraction of the limbs and overall greater locomotive efficiency (Foss, 2001). Of course, such a trait was not only found in entelodonts but in artiodactyls as a whole, likely being a response to predatory pressures from incumbent predatory clades arising at the same time as the artiodactyls (Foss, 2001). However, in the case of the entelodonts, such adaptations were not used for merely escaping predators. Rather, they were used to for another, much more lethal effect… Such notions are further reinforced by the entelodonts most formidable aspect, none either than their fearsome jaws, and in this respect, Archaeotherium excelled. Both for its size and in general, the head of Archaeotherium was massive, measuring 40-50 cm (1.3-1.6 ft) in length among average A. mortoni specimens, to up to 78 cm (~2.6 ft) in the larger “Megachoerus” specimens (Joeckel, 1990). Such massive skulls were supported and supplemented by equally massive neck muscles and ligaments, which attached to massive neural spines on the anterior thoracic vertebrae akin to a bisons hump as well as to the sternum, allowing Archaeotherium to keep its head aloft despite the skulls massive size (Effinger, 1998). Of course, with such a massive skull, it should come as no surprise that such skulls housed exceptionally formidable jaws as well, and indeed, the bite of Archaeotherium was an especially deadly one. Its zygomatic arches (cheek-bones) and its temporal fossa were enlarged and expanded, indicative of massive temporalis muscles that afforded Archaeotherium astoundingly powerful bites (Joeckel, 1990). This is further augmented by Archaeotherium’s massive jugal flanges (bony projections of the cheek), which supported powerful masseter muscles which enhanced chewing and mastication, as well as an enlarged postorbital bar that reinforced the skull against torsional stresses (Foss, 2001). Last but not least, powerful jaws are supplemented by an enlarged gape, facilitated by a low coronoid process and enlarged posterior mandibular tubercles (bony projections originating from the lower jaw), which provided an insertion site for sternum-to-mandible jaw abduction muscles, allowing for a more forceful opening of the jaw (Foss, 2001). All together, such traits suggest a massive and incredibly fearsome bite, perhaps the most formidable of any animal in its environment. Of course, none of such traits are especially indicative of a predatory lifestyle. Indeed, many modern non-predatory ungulates, like hippos, pigs and peccaries, also possess large, formidable skulls and jaws. However, in peeling back the layers, it is found there was more to the skull of Archaeotherium that lies in store. Indeed, when inspecting the animal closely, a unique mosaic of features is revealed; traits that make it out to be much more lethal than the average artiodactyl. On one hand, Archaeotherium possessed many traits similar to those of herbivores animals, as is expected of ungulates. For instance, its jaw musculature that allowed the lower jaw of Archaeotherium a full side-to-side chewing motion as in herbivores (whereas most carnivores can only move their lower jaw up and down)(Effinger, 1998). On the other hand, Archaeotherium wielded many other traits far more lethal in their morphology, less akin to a herbivore and far more akin to a bonafide predator. For instance, the aforementioned enlarged gape of Archaeotherium is a bizarre trait on a supposed herbivore, as such animals do not need large gapes to eat vegetation and thus have smaller, more restricted gapes. Conversely, many predatory lineages have comparatively large gapes, as larger gapes allow for the the jaws to grab on to more effectively larger objects, namely large prey animals (Joeckel, 1990). Such a juxtaposition, however, is most evident when discussing the real killing instruments of Archaeotherium — the teeth. More so than any facet of this animal, the teeth of Archaeotherium are the real stars of the show, showing both how alike it was compared to its herbivores counterparts and more importantly, how it couldn’t be more different. For instance, the molars of Archaeotherium were quite similar to modern herbivores ungulates, in that they were robust, bunodont, and were designed for crushing and grinding, similar in form and function to modern ungulates like peccaries (Joeckel, 1990). However, while the molars give the impression that Archaeotherium was a herbivore, the other teeth tell a very different story. The incisors, for example, were enlarged, sharpened, and fully interlocked (as opposed to the flat-topped incisors seen in herbivores ungulates), creating an incisor array that was seemingly ill-suited for cropping vegetation and much more adept at for gripping, puncturing and cutting (Joeckel, 1990). Even more formidable were the canines. Like the modern pigs from which entelodonts derived their nicknames, the canines of Archaeotherium were sharp and enlarged to form prominent tusk-like teeth, but unlike pigs, they were rounded in cross-section (similar to modern carnivores like big cats, indicating more durable canines that can absorb and resist torsional forces, such as those from struggling prey) and were serrated to form a distinct cutting edge (Effinger, 1998; Joeckel, 1990; Ruff & Van Valkenburgh, 1987). These canines, along with the incisors, interlock to stabilize the jaws while biting and dismantling in a carnivore-like fashion. More strikingly, the canines also seem to act as “occlusal guides,” wherein the canines help align the movement and position of the rear teeth as they come together, allowing for a more efficient shearing action by the rear teeth. This function is seen most prevalently modern carnivores mammals, and is evidenced by the canine tooth-wear, which is also analogous to modern predators like bears and canids (Joeckel, 1990). Indeed, going off such teeth alone, it is clear that Archaeotherium is far more predatory than expected of an ungulate. However, the real stars of the show, the teeth that truly betray the predatory nature of these ungulates, are the premolars. Perhaps the most carnivore-like teeth in the entelodont’s entire tooth row, the premolars of Archaeotherium, particularly the anterior premolars, are laterally compressed, somewhat conical in shape, and are weakly serrated to bear a cutting edge, giving them a somewhat carnivorous form and function of shearing and slicing (Effinger, 1998). Most strikingly of all, the premolars of Archaeotherium bear unique features similar not to modern herbivores, but to durophagous carnivores like hyenas, particularly apical wear patterns, highly thickened enamel, “zigzag-shaped” enamel prism layers (Hunter-Schraeger bands) on the premolars which is also seen in osteophagous animals like hyenas, and an interlocking premolar interface wherein linear objects (such as bones) inserted into jaws from the side would be pinned between the premolars and crushed (Foss, 2001). Taken together, these features do not suggest a diet of grass or vegetation like other ungulates. Rather, they suggest a far more violent diet, one including flesh as well as hard, durable foods, particularly bone. All in all, the evidence is clear. Archaeotherium and other entelodonts, unlike the rest of their artiodactyl kin, were not the passive herbivores as we envision ungulates today. Rather, they were willing, unrepentant meat-eaters that had a taste for flesh as well as foliage. Of course, even with such lines of evidence, its hard to conclude that Archaeotherium was a true predator. After all, its wide gape and durophagous teeth could have just as easily been used for scavenging or even to eat tough plant matter such as seeds or nuts, as in peccaries and pigs, which themselves share many of the same adaptations as Archaeotherium, include the more carnivorous ones (e.g. the wide gape, using the canines as an occlusal guide, etc.). How exactly do we know that these things were veritable predators and not pretenders to the title. To this end, there is yet one last piece of evidence, one that puts on full display the predatory prowess of Archaeotherium —evidence of a kill itself. Found within oligocene-aged sediment in what is now Wyoming, a collection of various fossil remains was found, each belonging to the ancient sheep-sized camel Poebrotherium, with many of the skeletal remains being disarticulated and even missing whole hindlimbs or even entire rear halves of their body. Tellingly, many of the remains bear extensive bite marks and puncture wounds across their surface. Upon close examination, the spacing and size of the punctures leave only one culprit: Archaeotherium. Of course, such an event could still have been scavenging; the entelodonts were consuming the remains of already dead, decomposed camels, explaining the bite marks. What was far more telling, however, was where the bite marks were found. In addition bite marks being found on the torso and lumbar regions of the camels, various puncture wounds were found on the skull and neck, which were otherwise uneaten. Scavengers rarely feast on the head to begin with; there is very little worthwhile meat on it besides the brain, cheek-muscles and eyes, and even if they did feed on the skull and neck, they would still eat it wholesale, not merely bite it and then leave it otherwise untouched. Indeed, it was clear that this was no mere scavenging event. Rather than merely consuming these camels, Archaeotherium was actively preying upon and killing them, dispatching them via a crushing bite to the skull or neck before dismembering and even bisecting the hapless camels with their powerful jaws to preferentially feast on their hindquarters (likely by swallowing the hindquarters whole, as the pelvis of Poebrotherium was coincidentally the perfect width for Archaeotherium to devour whole), eventually discarding the leftovers in meat caches for later consumption (Sundell, 1999). With this finding, such a feat of brutality leaves no doubt in ones mind as to what the true nature of Archaeotherium was. This was no herbivore, nor was it a simple scavenger. This was an active, rapacious predator, the most powerful in its entire ecosystem. Indeed, with such brutal evidence of predation frozen in time, combined with various dental, cranial, and post cranial adaptations of this formidable animal, it’s possible to paint a picture of how this formidable creature lived. Though an omnivore by trade, willing and able to feast on plant matter such as grass, roots and tubers, Archaeotherium was also a wanton predator that took just about any prey it wanted. Upon detecting its prey, it approached its vicim from ambush before launching itself at blazing speed. From there, its cursorial, hoofed legs, used by other ungulates for escape predation, were here employed to capture prey, carrying it at great speeds as it caught up to its quarry. Having closed the distance with its target, it was then that the entelodont brought its jaws to bear, grabbing hold of the victim with powerful jaws and gripping teeth to bring it to a screeching halt. If the victim is lucky, Archaeotherium will then kill it quickly with a crushing bite to the skull or neck, puncturing the brain or spinal cord and killing its target instantly. If not, the victim is eaten alive, torn apart while it’s still kicking, as modern boars will do today. In any case, incapacitated prey are subsequently dismantled, with the entelodont using its entire head and heavily-muscled necks to bite into and pull apart its victim in devastating “puncture-and pull’ bites (Foss, 2001). Prey would then finally be consumed starting at the hindquarters, with not even the bones of its prey being spared. Such brutality, though far from clean, drove home a singular truth: that during this time, ungulates were not just prey, that they were not the mere “predator-fodder” we know them as today. rather, they themselves were the predators themselves, dominating as superb hunters within their domain and even suppressing clades we know as predators today, least of all the carnivorans. Indeed, during this point in time, the age of the carnivorous ungulates had hit their stride, and more specifically, the age of entelodonts had begun. Of course, more so than any other ettelodont, Archaeotherium took to this new age with gusto. Archaeotherium lived from 35-28 million years ago during the late Eocene and early Oligocene in a locality known today as the White River Badlands, a fossil locality nestled along the Great Plains and Rocky Mountains. Though a chalky, barren landscape today, during the time of Archaeotherium, the White River Badlands was a swamp-like floodplain crisscrossed with rivers and interspersed with by a mosaic of forests concentrated around waterways, open woodlands and open plains. As with most ecosystems with such a lush disposition, this locale teemed with life, with ancient hornless rhinos, small horse-like hyracodonts and early camels roaming the open habitats while giant brontotheres, small early horses and strange, sheep-like ungulates called merycoidodonts (also known as “oreodonts”) dwelled within the dense forests. Within this locale, Archaeotherium stalked the open woodlands and riparian forests of its domain. Here, it acted as a dominant predator and scavenger across is territory, filling a niche similar to modern grizzly bears but far more predatory. Among its preferred food items would be plant matter such as roots, foliage and nuts, but also meat in the form of carrion or freshly caught prey. In this respect, smaller ungulates such as the fleet-footed camel Poebrotherium, a known prey item of Archaeotherium, would have made a for choice prey, as its small size would make it easy for Archaeotherium to dispatch with its powerful jaws, while the entelodonts swift legs gave it the speed necessary to keep pace with its agile prey. However, the entelodont didn’t have such a feast all to itself. Just as the badlands teemed with herbivores, so too did it teem with rival predators. Among their ranks included fearsome predators such as Hyaenodon, a powerful, vaguely dog-like predator up to the size of wolves (as in H. horridus) or even lions (as in the Eocene-aged H. megaloides, which was replaced by H. horridus during the Oligocene). Armed with a massive head, fierce jaws and a set of knife-like teeth that could cut down even large prey in seconds, these were some of the most formidable predators on the landscape. There were also the nimravids, cat-like carnivorans that bore saber-teeth to kill large prey in seconds, and included the likes of the lynx-sized Dinictis, the leopard-sized Hoplophoneus and even the jaguar-sized Eusmilus. Furthermore, there were amphicyonids, better known as the bear-dogs. Though known from much larger forms later on in their existence, during the late Eocene and Oligocene, they were much smaller and acted as the “canid-analogues” of the ecosystem, filling a role similar to wolves or coyotes. Last but not least, there were the bathornithid birds, huge cariamiform birds related to modern seriemas but much larger, which filled a niche similar to modern seriemas or secretary birds, albeit on a much larger scale. Given such competition, it would seem that Archaeotherium would have its hands full. However, things are not as they appear. For starters, habitat differences would mitigate high amounts of competition, as both Hyaenodon and the various nimravids occupy more specialized ecological roles (being a plains-specialist and forest-specialist, respectively) than did Archaeotherium, providing a buffer to stave off competition: More importantly, however, none of the aforementioned predators were simply big enough to take Archaeotherium on. During the roughly 7 million years existence of Archaeotherium, the only carnivore that matched it in size was H. megaloides, and even that would have an only applied to average A. mortoni individuals, not to the much larger, bison-sized “Megachoerus” individuals. The next largest predator at that point would be the jaguars-sized Eusmilus (specifically E. adelos) which would have only been a bit more than half the size of even an average A. mortoni. Besides that, virtually every other predator on the landscape was simply outclassed by the much larger entelodont in terms of size and brute strength. As such, within its domain, Archaeotherium had total, unquestioned authority, dominating the other predators in the landscape and likely stealing their kills as well. In fact, just about the only threat Archaeotherium had was other Archaeotherium, as fossil bite marks suggest that this animal regularly and fraglantly engaged in intraspecific combat, usually through face-biting and possibly even jaw-wrestling (Effinger, 1998; Tanke & Currie, 1998). Nevertheless, it was clear that Archaeotherium was the undisputed king of the badlands; in a landscape of hyaenodonts and carnivorans galore, it was a hoofed ungulate that reigned supreme. However, such a reign would not last. As the Eocene transitioned into the Eocene, the planet underwent an abrupt cooling and drying phase known as Eocene-Oligocene Transition or more simply the Grande Coupure. This change in climate would eliminate the sprawling wetlands and river systems that Archaeotherium had been depending on, gradually replacing it with drier and more open habitats. To its credit, Archaeotherium did manage to hang on, persisting well after the Grand-Coupure had taken place, but in the end the damage had been done; Archaeotherium was a dead-man-walking. Eventually, by around 28 million years ago, Archaeotherium would go extinct, perishing due to this change in global climate (Gillham, 2019). Entelodonts as a whole would persist into the Miocene, producing some of their largest forms ever known in the form of the bison-sized Daeodon (which was itself even more carnivorous than Archaeotherium), however they too would meet the same fate as their earlier cousins. By around 15-20 million years ago, entelodonts as a whole would go extinct. However, while the entelodonts may have perished, this was not the end of carnivorous ungulates as a whole. Recall that the cetacodontamorphs, the lineage of artiodactyls that produced the entelodonts, left behind two living descendants. The first among them were the hippos, themselves fairly frequent herbivores. The second of such lineage, however, was a different story. Emerging out of South Asia, this lineage of piscivorous cetacodontamorphs, in a an attempt to further specialize for the fish-hunting lifestyle, began to delve further and further into the water, becoming more and more aquatic and the millennia passed by. At a certain point, these carnivorous artiodactlys had become something completely unrecognizable from their original hoofed forms. Their skin became hairless and their bodies became streamlined for life in water. Their hoofed limbs grew into giant flippers for steering in the water and their previously tiny tails became massive and sported giant tail flukes for aquatic propulsion. Their noses even moved to the tip of their head, becoming a blowhole that would be signature to this clade as a whole. Indeed, this clade was none other than the modern whales, themselves derived, carnivorous ungulates that had specialized for a life in the water, and in doing so, became the some of the most dominant aquatic predators across the globe for millions of years. Indeed, though long gone, the legacy of the entelodonts and of predatory ungulates as a whole, a legacy Archaeotherium itself had helped foster, lives on in these paragons of predatory prowess, showing that the ungulates are more than just the mere “prey” that they are often made out to be. Moreover, given the success that carnivorous ungulates had enjoyed in the past and given how modern omnivorous ungulates like boar dabble in predation themselves, perhaps, in the distant future, this planet may see the rise of carnivorous ungulates once again, following in the footsteps left behind by Archaeotherium and the other predatory ungulates all those millions of years ago. |
2024.05.14 16:19 Hiebster The Courage to Use Your Own Understanding
According to a new report from the UK government, one of the biggest threats to social cohesion is "freedom-restricting harassment." Average people are more familiar with the term "cancel culture" and are well aware of its destructive effects. Still, they are generally discouraged from mentioning it in the interests of diversity and inclusion, despite this being common sense. The courage to use our own understanding often puts us at odds with so-called "conventional wisdom" and that's what this post is about. submitted by Hiebster to BreakingTheNarrative [link] [comments] |
2024.05.14 16:18 Hiebster The Courage to Use Your Own Understanding
According to a new report from the UK government, one of the biggest threats to social cohesion is "freedom-restricting harassment." Average people are more familiar with the term "cancel culture" and are well aware of its destructive effects. Still, they are generally discouraged from mentioning it in the interests of diversity and inclusion, despite this being common sense. The courage to use our own understanding often puts us at odds with so-called "conventional wisdom" and that's what this post is about. submitted by Hiebster to substackpostmedium [link] [comments] |
2024.05.14 16:18 Hiebster The Courage to Use Your Own Understanding
According to a new report from the UK government, one of the biggest threats to social cohesion is "freedom-restricting harassment." Average people are more familiar with the term "cancel culture" and are well aware of its destructive effects. Still, they are generally discouraged from mentioning it in the interests of diversity and inclusion, despite this being common sense. The courage to use our own understanding often puts us at odds with so-called "conventional wisdom" and that's what this post is about. submitted by Hiebster to JordanPeterson [link] [comments] |
2024.05.14 16:17 Hiebster The Courage to Use Your Own Understanding
According to a new report from the UK government, one of the biggest threats to social cohesion is "freedom-restricting harassment." Average people are more familiar with the term "cancel culture" and are well aware of its destructive effects. Still, they are generally discouraged from mentioning it in the interests of diversity and inclusion, despite this being common sense. The courage to use our own understanding often puts us at odds with so-called "conventional wisdom" and that's what this post is about. submitted by Hiebster to CanadianConservative [link] [comments] |
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2024.05.14 16:15 jnordwick Calling my season officially over
2024.05.14 16:15 AutoModerator [Tuesday] Daily Music Discussion - 14 May 2024
2024.05.14 16:12 gabrieu Advices for chubby guys?